Gnaphosidae: Drassodes

In case you find an error or have a specific suggestion, please follow this link: click
1
Males

  2  

- Females

  8  

2
(1)
Tibial apophysis of pedipalp undivided

  3  

speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1976a)
- Tibial apophysis of pedipalp forked

  6  

3
(2)
Distal bulbal apophysis small, embolus not covered

  4  

speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1976a)
-

world

add

cp

Distal bulbal apophysis large, partly covers embolus

Drassodes pubescens (Thorell, 1856)

female

speciesimage
male Pedipalp
(Azheganova 1968)
speciesimage
male Pedipalp, ventral view
(Braendegaard 1966)
speciesimage
male Pedipalp, ventral view
(Miller 1971)
speciesimage
male Pedipalp, ventral view
(Noordam 1992)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Noordam 1992)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
Chelicerae, ventral view
(Locket & Millidge 1951)
4
(3)

world

add

cp

Tibial apophysis with serrated margin; embolus small; distal apophysis hook-shaped

Drassodes villosus (Thorell, 1856)

female

speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
- Different

  5  

5
(4)

world

add

cp

Chelicerae with three teeth as fig.; tibial apophysis with smooth margin; embolus larger; distal apophysis not hook-shaped

Drassodes lapidosus (Walckenaer, 1802)

female

speciesimage
male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1976a)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1976a)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Chatzaki et al. 2002b)
speciesimage
male Chelicerae, ventral view
(Roberts 1995)
speciesimage
Chelicerae, ventral view
(Locket & Millidge 1951)
-

world

add

cp

Chelicerae with 3 teeth as fig.

Drassodes cupreus (Blackwall, 1834)

female

speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Chelicerae, ventral view
(Roberts 1995)
speciesimage
Chelicerae, ventral view
(Locket & Millidge 1951)
speciesimage
Chelicerae, ventral view
(Locket & Millidge 1951)
6
(2)

world

add

cp

Embolus with striking distal apophysis

Drassodex hypocrita (Simon, 1878)

female

speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp
(Roberts 1998)
- Embolus without distal apophysis

  7  

7
(6)

world

add

cp

Last embolic loop > 1/2 width of bulbus

Drassodex heeri (Pavesi, 1873)

female

speciesimage
male Pedipalp
(Wiehle 1967)
speciesimage
male Pedipalp, ventral view
(Wiehle 1967)
-

world

add

cp

Last embolic loop ≤ 1/2 width of bulbus

Drassodex hispanus (L. Koch, 1866)

female

8
(1)
Epigyne with deep groove, sclerotised area not reaching epigastric furrow

  9  

speciesimage
female Epigyne
(Wiehle 1967)
- Epigyne without groove, sclerotised area reaching epigastric furrow

  11  

9
(8)
Entrance duct spirally looping around wide tubes of copulatory opening

  10  

speciesimage
female Vulva
(Heimer & Nentwig 1991)
-

world

add

cp

Entrance duct different; epigyne as fig.

Drassodex heeri (Pavesi, 1873)

male

speciesimage
female Epigyne
(Wiehle 1967)
speciesimage
female Vulva
(Wiehle 1967)
10
(9)

world

add

cp

Entrance duct with 2 loops; epigyne as fig.

Drassodex hypocrita (Simon, 1878)

male

speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
-

world

add

cp

Entrance duct with more than 3 loops; epigyne as fig.

Drassodex hispanus (L. Koch, 1866)

male

speciesimage
female Epigyne
(Hervé et al. 2009)
speciesimage
female Vulva, dorsal view
(Hervé et al. 2009)
11
(8)
Epigyne with lateral bars

  12  

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Epigyne
(Roberts 1995)
-

world

add

cp

Different; epigyne and vulva as fig.

Parasyrisca vinosa (Simon, 1878)

speciesimage
female Epigyne (courtesy of the AMNH)
(Ovtsharenko et al. 1995)
speciesimage
female Vulva (courtesy of the AMNH)
(Ovtsharenko et al. 1995)
12
(11)

world

add

cp

Epigynal ridges almost touching each other in the middle part

Drassodes pubescens (Thorell, 1856)

male

speciesimage
female Epigyne
(Azheganova 1968)
speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Noordam 1992)
speciesimage
female Epigyne
(Pérez-Pérez 1985)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Guiming et al. 2002)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Vulva, dorsal view
(Tullgren 1946)
speciesimage
female Vulva
(Pérez-Pérez 1985)
speciesimage
female Vulva
(Guiming et al. 2002)
- Different

  13  

13
(12)

world

add

cp

Central epigynal area trapezoid, wider than long

Drassodes villosus (Thorell, 1856)

male

speciesimage
female Epigyne
(Azheganova 1968)
speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
speciesimage
female Vulva, dorsal view
(Tullgren 1946)
- Central epigynal area roundish, longer than wide

  14  

14
(13)

world

add

cp

Epigyne and vulva as figs.

Drassodes lapidosus (Walckenaer, 1802)

male

speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1976a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Locket et al. 1974)
speciesimage
female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1976a)
speciesimage
female Vulva, dorsal view
(Tullgren 1946)
speciesimage
female Vulva
(Chatzaki et al. 2002b)
-

world

add

cp

Epigyne very similar to lapidosus, but lateral parts a bit wider, median parts narrower

Drassodes cupreus (Blackwall, 1834)

male

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket et al. 1974)
Further taxa

world

add

cp

1. Drassodes albicans (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Simon 1914)
speciesimage
female Vulva
(Chatzaki et al. 2002b)
speciesimage
Chelicerae
(Chatzaki et al. 2002b)

world

add

cp

2. Drassodes andorranus Denis, 1938

speciesimage
female Epigyne, ventral view
(Denis 1938)

world

add

cp

3. Drassodes archibensis Ponomarev & Alieva, 2008

speciesimage
male Pedipalp, ventral view
(Ponomarev & Alieva 2008)
speciesimage
male Pedipalp, ventral view
(Ponomarev & Alieva 2008)

world

add

cp

4. Drassodes auritus Schenkel, 1963

speciesimage
male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2004)
speciesimage
female Epigyne, ventral view
(Ponomarev & Tsvetkov 2004)
speciesimage
female Epigyne, ventral view
(Schenkel 1963)
speciesimage
female Vulva, dorsal view
(Ponomarev & Tsvetkov 2004)

world

add

cp

5. Drassodes bifidus Kovblyuk & Seyyar, 2009

speciesimage
male Pedipalp, retrolateral view
(Kovblyuk et al. 2009)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2009)
speciesimage
female Epigyne/vulva, dorsal view
(Seyyar et al. 2015)
speciesimage
female Epigyne/vulva, ventral view
(Seyyar et al. 2015)

world

add

cp

6. Drassodes canaglensis Caporiacco, 1927

speciesimage
female Epigyne
(Caporiacco 1927)

world

add

cp

7. Drassodes caspius Ponomarev & Tsvetkov, 2006

speciesimage
male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2006)
speciesimage
male Chelicera, posterior view
(Ponomarev & Tsvetkov 2006)
speciesimage
female Epigyne, ventral view
(Ponomarev & Tsvetkov 2006)

world

add

cp

8. Drassodes chybyndensis Esyunin & Tuneva, 2002

speciesimage
male Pedipalp, dorsal view
(Esyunin & Tuneva 2002)
speciesimage
male Pedipalp, ventral view
(Esyunin & Tuneva 2002)
speciesimage
male Pedipalp, retrolateral view
(Esyunin & Tuneva 2002)
speciesimage
male Cheliceral teeth, anterior view
(Esyunin & Tuneva 2002)
speciesimage
female Epigyne, ventral view
(Esyunin & Tuneva 2002)
speciesimage
female Vulva
(Esyunin & Tuneva 2002)
speciesimage
female Cheliceral teeth, anterior view
(Esyunin & Tuneva 2002)

world

add

cp

9. Drassodes dagestanus Ponomarev & Alieva, 2008

speciesimage
male Pedipalp, ventral view
(Ponomarev & Alieva 2008)
speciesimage
male Pedipalp, ventral view
(Ponomarev & Alieva 2008)
speciesimage
female Epigyne, ventral view
(Ponomarev & Alieva 2008)
speciesimage
female Vulva, dorsal view
(Ponomarev & Alieva 2008)

world

add

cp

10. Drassodes difficilis (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Hervé & Rollard 2009)
speciesimage
male Pedipalp, retrolateral view
(Hervé & Rollard 2009)
speciesimage
male Pedipalp, ventral view
(Alderweireldt & De Boever 2016)
speciesimage
male Pedipalp, retrolateral view
(Alderweireldt & De Boever 2016)
speciesimage
male Chelicera, ventral view
(Hervé & Rollard 2009)
speciesimage
female Epigyne, ventral view
(Alderweireldt & De Boever 2016)
speciesimage
female Vulva, ventral view
(Alderweireldt & De Boever 2016)
speciesimage
female Vulva, ventral view
(Alderweireldt & De Boever 2016)

world

add

cp

11. Drassodes fugax (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Hervé & Rollard 2009)
speciesimage
male Pedipalp, retrolateral view
(Hervé & Rollard 2009)
speciesimage
male Tibial apophysis
(Simon 1914)
speciesimage
female Epigyne
(Simon 1914)
speciesimage
female Epigyne, ventral view
(Hervé & Rollard 2009)
speciesimage
female Vulva, dorsal view
(Hervé & Rollard 2009)

world

add

cp

12. Drassodes inermis (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Hervé & Rollard 2009)
speciesimage
male Pedipalp, retrolateral view
(Hervé & Rollard 2009)
speciesimage
male Chelicera, ventral view
(Hervé & Rollard 2009)
speciesimage
female Epigyne
(Simon 1914)
speciesimage
female Epigyne, ventral view
(Hervé & Rollard 2009)
speciesimage
female Vulva, dorsal view
(Hervé & Rollard 2009)
speciesimage
female Chelicera, ventral view
(Hervé & Rollard 2009)

world

add

cp

13. Drassodes lacertosus (O. P.-Cambridge, 1872)

speciesimage
male Pedipalp, retrolateral view
(Levy 2004)
speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Palpal femur
(Levy 2004)
speciesimage
male Palpal tibia, retrolateral view
(Levy 2004)
speciesimage
male Chelicera, ventral view
(Levy 2004)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Vulva, dorsal view
(Levy 2004)

world

add

cp

14. Drassodes lapidosus bidens (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Simon 1914)
speciesimage
male Cheliceral teeth
(Simon 1914)

world

add

cp

15. Drassodes luteomicans (Simon, 1878)

speciesimage
male Pedipalp, retrolateral view
(Pantini et al. 2013)
speciesimage
male Pedipalp, ventral view
(Pantini et al. 2013)
speciesimage
female Epigyne
(Simon 1914)
speciesimage
female Epigyne, ventral view
(Pantini et al. 2013)
speciesimage
female Vulva, ventral view
(Pantini et al. 2013)
speciesimage
Chelicera, posterior view
(Pantini et al. 2013)

world

add

cp

16. Drassodes lutescens (C. L. Koch, 1839)

speciesimage
male Pedipalp, retrolateral view
(Levy 2004)
speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp, ventral view
(Di Franco 1994a)
speciesimage
male Pedipalp
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp
(Di Franco 1994a)
speciesimage
male Palpal tibia
(Levy 2004)
speciesimage
male Chelicera, ventral view
(Levy 2004)
speciesimage
female Epigyne
(Di Franco 1994a)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Di Franco 1994a)
speciesimage
female Epigyne
(Crespo et al. 2009b)
speciesimage
female Vulva
(Di Franco 1994a)
speciesimage
female Vulva, dorsal view
(Levy 2004)
speciesimage
female Vulva
(Chatzaki et al. 2002b)
speciesimage
female Vulva
(Di Franco 1994a)

world

add

cp

17. Drassodes mandibularis (L. Koch, 1866)

speciesimage
female Epigyne, ventral view
(L Koch 1866)

world

add

cp

18. Drassodes montenegrinus (Kulczyński, 1897)

speciesimage
female Epigyne/vulva
(Chyzer & Kulczyński 1897)
speciesimage
female Epigyne
(Chyzer & Kulczyński 1897)

world

add

cp

19. Drassodes natali Esyunin & Tuneva, 2002

speciesimage
male Pedipalp, ventral view
(Esyunin & Tuneva 2002)
speciesimage
male Pedipalp, dorsal view
(Esyunin & Tuneva 2002)
speciesimage
male Palpal tibia, ventral view
(Esyunin & Tuneva 2002)
speciesimage
male Chelicera, anterior view
(Esyunin & Tuneva 2002)
speciesimage
female Epigyne, ventral view
(Esyunin & Tuneva 2002)
speciesimage
female Vulva
(Esyunin & Tuneva 2002)
speciesimage
female Chelicera, anterior view
(Esyunin & Tuneva 2002)

world

add

cp

20. Drassodes orientalis (L. Koch, 1866)

speciesimage
female Epigyne
(L Koch 1866)

world

add

cp

21. Drassodes placidulus Simon, 1914

speciesimage
male Pedipalp, ventral view
(Simon 1914)

world

add

cp

22. Drassodes platnicki Song, Zhu & Zhang, 2004

speciesimage
male Pedipalp, ventral view
(Esyunin & Tuneva 2002)
speciesimage
male Pedipalp, ventral view
(Song et al. 1999)
speciesimage
female Epigyne, ventral view
(Esyunin & Tuneva 2002)
speciesimage
female Epigyne, ventral view
(Song et al. 1999)
speciesimage
female Vulva
(Esyunin & Tuneva 2002)

world

add

cp

23. Drassodes rostratus Esyunin & Tuneva, 2002

speciesimage
male Pedipalp, dorsal view
(Esyunin & Tuneva 2002)
speciesimage
male Pedipalp, ventral view
(Esyunin & Tuneva 2002)
speciesimage
male Palpal tibia, lateral view
(Esyunin & Tuneva 2002)
speciesimage
male Chelicera, frontal view
(Esyunin & Tuneva 2002)
speciesimage
male Chelicera, lateral view
(Esyunin & Tuneva 2002)
speciesimage
female Epigyne
(Esyunin et al. 2007)

world

add

cp

24. Drassodes rubidus (Simon, 1878)

speciesimage
male Pedipalp, ventral view
(Simon 1914)
speciesimage
female Epigyne
(Simon 1914)

world

add

cp

25. Drassodes serratichelis (Roewer, 1928)

speciesimage
male Pedipalp
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Pedipalp, retrolateral view
(Levy 2004)
speciesimage
male Pedipalp, prolateral view
(Wunderlich 2011a)
speciesimage
male Pedipalp, ventral view
(Wunderlich 2011a)
speciesimage
male Embolic division
(Wunderlich 2011a)
speciesimage
male Prosoma, dorsal view
(Wunderlich 2011a)
speciesimage
male Trochanter IV, ventral view
(Wunderlich 2011a)
speciesimage
male Metatarsus III, ventral view
(Wunderlich 2011a)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Vulva
(Chatzaki et al. 2002b)
speciesimage
female Vulva, dorsal view
(Levy 2004)

world

add

cp

26. Drassodes striatus (L. Koch, 1866)

speciesimage
female Epigyne
(L Koch 1866)

world

add

cp

27. Drassodes thaleri Hervé & Rollard, 2009

speciesimage
male Pedipalp, ventral view
(Hervé & Rollard 2009)
speciesimage
male Pedipalp, retrolateral view
(Hervé & Rollard 2009)
speciesimage
male Chelicera, ventral view
(Hervé & Rollard 2009)
speciesimage
female Epigyne, ventral view
(Hervé & Rollard 2009)
speciesimage
female Vulva, dorsal view
(Hervé & Rollard 2009)
speciesimage
female Chelicera, ventral view
(Hervé & Rollard 2009)

world

add

cp

28. Drassodes unicolor (O. P.-Cambridge, 1872)

speciesimage
male Pedipalp
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002b)
speciesimage
male Pedipalp, retrolateral view
(Levy 2004)
speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Tibial apophysis
(Levy 2004)
speciesimage
male Chelicera, ventral view
(Levy 2004)
speciesimage
female Epigyne
(Chatzaki et al. 2002b)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Vulva, dorsal view
(Levy 2004)
speciesimage
female Chelicera, ventral view
(Levy 2004)
29. Drassodes charcoviae (Thorell, 1875) world
30. Drassodes paroculus Simon, 1893 world
31. Drassodes rhodanicus Simon, 1914 world
32. Drassodes similis Nosek, 1905 world