Gnaphosidae: Haplodrassus

In case you find an error or have a specific suggestion, please follow this link: click
1
Males

  2  

- Females

  12  

2
(1)

world

add

cp

Tibial apophysis narrow; body length smaller or equal 4.0 mm; prosoma length smaller or equal 2.0 mm

Haplodrassus minor (O. P.-Cambridge, 1879)

female

speciesimage
male Pedipalp, ventral view
(Locket et al. 1974)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Palpal tibia
(Locket et al. 1974)
speciesimage
male Tibial apophysis
(Miller & Buchar 1977)
- Tibial apophysis different and/or body length larger

  3  

3
(2)
Tibial apophysis distally broadened

  4  

- Tibial apophysis distally not broadened

  8  

4
(3)

world

add

cp

Distal bulbal apophysis with strikingly serrated margin

Haplodrassus kulczynskii Lohmander, 1942

female

speciesimage
male Pedipalp, ventral view
(Miller 1971)
speciesimage
male Pedipalp
(Roberts 1998)
speciesimage
male Tibial apophysis
(Miller 1971)
- Distal bulbal apophysis different

  5  

5
(4)
Distal bulbal apophysis long and slender

  6  

- Distal bulbal apophysis broader and

  9  

6
(5)

world

add

cp

Pedipalp as fig.; tibial apophysis distally hardly broadened

Haplodrassus bohemicus Miller & Buchar, 1977

female

speciesimage
male Pedipalp, ventral view
(Miller & Buchar 1977)
speciesimage
male Embolus
(Miller & Buchar 1977)
speciesimage
male Tibial apophysis
(Miller & Buchar 1977)
- Different

  7  

7
(6)

world

add

cp

Distal bulbal apophysis distinctly keeled

Haplodrassus signifer (C. L. Koch, 1839)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1966)
speciesimage
male Pedipalp, ventral view
(Noordam 1992)
speciesimage
male Pedipalp
(Noordam 1992)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Tip of pedipalp
(Locket & Millidge 1951)
speciesimage
male Tibial apophysis
(Braendegaard 1966)
speciesimage
male Tibial apophysis
(Noordam 1992)
speciesimage
male Tibial apophysis
(Locket & Millidge 1951)
-

world

add

cp

Distal bulbal apophysis without keel

Haplodrassus aenus Thaler, 1984

female

speciesimage
male Pedipalp, ventral view
(Thaler 1984a)
speciesimage
male Tibial apophysis
(Thaler 1984a)
8
(3)

world

add

cp

Plate-like bulge of distal bulbal apophysis situated distally, overtops tip of apophysis; tibial apophysis distinctly broadened distally

Haplodrassus soerenseni (Strand, 1900)

female

speciesimage
male Pedipalp, ventral view
(Miller 1971)
speciesimage
male Pedipalp, ventral view
(Locket et al. 1974)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Locket & Millidge 1953)
speciesimage
male Embolic division, ventral view
(Locket & Millidge 1953)
speciesimage
male Tibial apophysis
(Miller & Buchar 1977)
-

world

add

cp

Plate-like bulge of distal bulbal apophysis situated basally; tibia apophysis only little broadened

Haplodrassus umbratilis (L. Koch, 1866)

female

speciesimage
male Pedipalp, ventral view
(Miller 1971)
speciesimage
male Pedipalp, ventral view
(Locket et al. 1974)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Palpal tibia
(Locket et al. 1974)
speciesimage
male Tibial apophysis
(Miller & Buchar 1977)
9
(5)
Distal bulbal apophysis distinctly broadened distally

  10  

-

world

add

cp

Distal bulbal apophysis not broadened distally and with 2 tooth-like structures

Haplodrassus dalmatensis (L. Koch, 1866)

female

speciesimage
male Pedipalp, ventral view
(Noordam 1992)
speciesimage
male Pedipalp, ventral view
(Locket et al. 1974)
speciesimage
male Pedipalp
(Noordam 1992)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Tibial apophysis
(Noordam 1992)
speciesimage
male Tibial apophysis
(Locket & Millidge 1951)
10
(9)

world

add

cp

Distal bulbal apophysis ± straight, basally with a tooth; tibial apophysis as fig.

Haplodrassus cognatus (Westring, 1861)

female

speciesimage
male Pedipalp, ventral view
(Miller & Buchar 1977)
speciesimage
male Tibial apophysis
(Miller & Buchar 1977)
- Distal bulbal apophysis ± rectangularly bent, without tooth

  11  

11
(10)

world

add

cp

Tibial apophysis large, reaching 1/2 of bulbal length, distinctly keeled

Haplodrassus silvestris (Blackwall, 1833)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1966)
speciesimage
male Pedipalp, ventral view
(Noordam 1992)
speciesimage
male Pedipalp
(Noordam 1992)
speciesimage
male Pedipalp
(Roberts 1995)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Tip of pedipalp
(Locket & Millidge 1951)
speciesimage
male Tibial apophysis
(Noordam 1992)
speciesimage
male Tibial apophysis
(Locket & Millidge 1951)
-

world

add

cp

Tibial apophysis small, reaching about 1/3 of bulbal length, without distinct keel

Haplodrassus moderatus (Kulczyński, 1897)

female

speciesimage
male Pedipalp, ventral view
(Miller 1971)
speciesimage
male Pedipalp
(Roberts 1998)
12
(1)
Epigyne with several transverse folds

  13  

- Different

  14  

13
(12)

world

add

cp

Bar of anterior epigynal margin narrow; depressed area between the sclerotised pockets narrow

Haplodrassus silvestris (Blackwall, 1833)

male

speciesimage
male Tip of pedipalp
(Locket & Millidge 1951)
speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Noordam 1992)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Miller & Buchar 1977)
speciesimage
female Vulva, ventral view
(Tullgren 1946)
-

world

add

cp

Bar of anterior epigynal margin narrow as broad as epigyne; depressed area between sclerotised pockets broad and with strong indentations

Haplodrassus cognatus (Westring, 1861)

male

speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Miller 1971)
speciesimage
female Vulva
(Miller & Buchar 1977)
speciesimage
female Vulva, ventral view
(Tullgren 1946)
14
(12)
Sclerotised epigynal pockets with shallow protrusion

  15  

- Sclerotised epigynal pockets without protrusion

  17  

15
(14)

world

add

cp

Protrusion very elongated

Haplodrassus kulczynskii Lohmander, 1942

male

speciesimage
female Epigyne
(Miller 1971)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva
(Miller & Buchar 1977)
- Different

  16  

16
(15)

world

add

cp

Protrusion slantingly forward-directed

Haplodrassus umbratilis (L. Koch, 1866)

male

speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket et al. 1974)
speciesimage
female Vulva
(Miller & Buchar 1977)
speciesimage
female Vulva, ventral view
(Tullgren 1946)
-

world

add

cp

Protrusions directed more towards median region

Haplodrassus soerenseni (Strand, 1900)

male

17
(14)
Margins of sclerotised, black epigynal pockets parallel or slightly roundish curved

  18  

- Margins of sclerotised, black epigynal pockets strongly converging or diverging posteriorly

  21  

18
(17)
Margins on the inner side roundish curved

  19  

- Margins on the inner side straight

  20  

19
(18)

world

add

cp

Epigyne and vulva as figs.

Haplodrassus signifer (C. L. Koch, 1839)

male

speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Noordam 1992)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1975)
speciesimage
female Vulva, ventral view
(Tullgren 1946)
-

world

add

cp

Epigyne and vulva as figs.

Haplodrassus bohemicus Miller & Buchar, 1977

male

speciesimage
female Epigyne
(Miller & Buchar 1977)
speciesimage
female Vulva
(Miller & Buchar 1977)
20
(18)

world

add

cp

Epigyne and vulva as figs.

Haplodrassus aenus Thaler, 1984

male

speciesimage
female Epigyne/vulva, ventral view
(Thaler 1984a)
speciesimage
female Epigyne/vulva, ventral view
(Thaler 1984a)
-

world

add

cp

Epigyne and vulva as figs.

Haplodrassus minor (O. P.-Cambridge, 1879)

male

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
21
(17)

world

add

cp

Margins diverging towards posterior

Haplodrassus dalmatensis (L. Koch, 1866)

male

speciesimage
female Epigyne
(Miller 1971)
speciesimage
female Epigyne
(Braendegaard 1966)
speciesimage
female Epigyne
(Miller 1971)
speciesimage
female Epigyne
(Noordam 1992)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Miller & Buchar 1977)
-

world

add

cp

Margins converging towards posterior

Haplodrassus moderatus (Kulczyński, 1897)

male

speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva, ventral view
(Tullgren 1946)
Further taxa

world

add

cp

1. Haplodrassus acrotirius (Roewer, 1928)

speciesimage
female Epigyne
(Chatzaki et al. 2002a)
speciesimage
female Vulva
(Chatzaki et al. 2002a)

world

add

cp

2. Haplodrassus caspius Ponomarev & Belosludtsev, 2008

speciesimage
male Pedipalp, retrolateral view
(Kovblyuk et al. 2013d)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2013d)
speciesimage
male Pedipalp, prolateral view
(Kovblyuk et al. 2013d)
speciesimage
male Bulbus, dorsal view
(Kovblyuk et al. 2013d)
speciesimage
male Bulbus, apical view
(Kovblyuk et al. 2013d)
speciesimage
female Epigyne, ventral view
(Ponomarev et al. 2008)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2013d)
speciesimage
female Vulva, dorsal view
(Ponomarev et al. 2008)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2013d)
speciesimage
female Eyes, frontal view
(Kovblyuk et al. 2013d)
speciesimage
female Chelicera, posterior view
(Kovblyuk et al. 2013d)

world

add

cp

3. Haplodrassus caucasius Ponomarev & Dvadnenko, 2013

speciesimage
male Pedipalp, ventral view
(Ponomarev & Dvadnenko 2013)
speciesimage
male Pedipalp, retrolateral view
(Ponomarev & Dvadnenko 2013)
speciesimage
female Epigyne, ventral view
(Ponomarev & Dvadnenko 2013)

world

add

cp

4. Haplodrassus cognatus ermolajewi Lohmander, 1942

speciesimage
female Epigyne, ventral view
(Ermolajev 1928)

world

add

cp

5. Haplodrassus concertor (Simon, 1878)

speciesimage
male Apical part of bulbus
(Simon 1914)
speciesimage
female Epigyne
(Simon 1914)

world

add

cp

6. Haplodrassus creticus (Roewer, 1928)

speciesimage
male Pedipalp, ventral view
(Chatzaki et al. 2002a)
speciesimage
male Pedipalp, retrolateral view
(Chatzaki et al. 2002a)
speciesimage
male Bulbus
(Chatzaki et al. 2002a)
speciesimage
male Palpal tibia, dorsal view
(Chatzaki et al. 2002a)
speciesimage
female Epigyne
(Chatzaki et al. 2002a)
speciesimage
female Vulva
(Chatzaki et al. 2002a)

world

add

cp

7. Haplodrassus dalmatensis pictus (Thorell, 1875)

speciesimage
male Pedipalp, ventral view
(Kulczyński 1899)

world

add

cp

8. Haplodrassus grazianoi Caporiacco, 1948

speciesimage
female Epigyne
(Caporiacco 1948)

world

add

cp

9. Haplodrassus hiemalis (Emerton, 1909)

speciesimage
male Pedipalp, ventral view
(Marusik & Logunov 1995b)
speciesimage
male Embolus, apical view
(Marusik & Logunov 1995b)
speciesimage
male Palpal tibia, retroapical view
(Marusik & Logunov 1995b)
speciesimage
female Epigyne, ventral view
(Marusik & Logunov 1995b)
speciesimage
female Vulva, dorsal view
(Marusik & Logunov 1995b)

world

add

cp

10. Haplodrassus ibericus Melic, Silva & Barrientos, 2016

speciesimage
male Pedipalp, ventro-apical view
(Melic et al. 2016)
speciesimage
male Pedipalp, prolateral view
(Melic et al. 2016)
speciesimage
male Pedipalp, retrolateral view
(Melic et al. 2016)
speciesimage
male Pedipalp, ventral view
(Melic et al. 2016)
speciesimage
male Tibial apophysis, dorsal view
(Melic et al. 2016)
speciesimage
female Epigyne
(Melic et al. 2016)
speciesimage
female Vulva
(Melic et al. 2016)

world

add

cp

11. Haplodrassus invalidus (O. P.-Cambridge, 1872)

speciesimage
male Pedipalp, ventral view
(Di Franco 1996)
speciesimage
male Pedipalp
(Di Franco 1996)
speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2012b)
speciesimage
male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage
male Tibial apophysis
(Levy 2004)
speciesimage
male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage
female Epigyne
(Di Franco 1996)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Vulva
(Di Franco 1996)
speciesimage
female Vulva, dorsal view
(Levy 2004)

world

add

cp

12. Haplodrassus isaevi Ponomarev & Tsvetkov, 2006

speciesimage
male Pedipalp, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2012b)
speciesimage
male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2006)
speciesimage
male Pedipalp, retrolateral view
(Ponomarev & Tsvetkov 2006)
speciesimage
male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2012b)
speciesimage
female Epigyne, ventral view
(Ponomarev & Tsvetkov 2006)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2012b)

world

add

cp

13. Haplodrassus ivlievi Ponomarev, 2015

speciesimage
female Epigyne, ventral view
(Ponomarev 2015)
speciesimage
female Vulva, dorsal view
(Ponomarev 2015)

world

add

cp

14. Haplodrassus macellinus (Thorell, 1871)

speciesimage
male Pedipalp, ventral view
(Simon 1878)
speciesimage
male Pedipalp, retrolateral view
(Lecigne unpubl.)
speciesimage
male Pedipalp, ventral view
(Lecigne unpubl.)
speciesimage
female Epigyne, ventral view
(Simon 1878)

world

add

cp

15. Haplodrassus macellinus hebes (O. P.-Cambridge, 1874)

speciesimage
male Palpal tibia
(O. Pickard-Cambridge 1874)

world

add

cp

16. Haplodrassus mediterraneus Levy, 2004

speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Palpal tibia, retrolateral view
(Levy 2004)
speciesimage
female Epigyne, ventral view
(Levy 2004)
speciesimage
female Vulva, dorsal view
(Levy 2004)

world

add

cp

17. Haplodrassus morosus (O. P.-Cambridge, 1872)

speciesimage
male Pedipalp, ventral view
(Levy 2004)
speciesimage
male Tibial apophysis
(Levy 2004)
speciesimage
female Epigyne
(Levy 2004)
speciesimage
female Vulva
(Levy 2004)

world

add

cp

18. Haplodrassus ovtchinnikovi Ponomarev, 2008

speciesimage
male Pedipalp, retrolateral view
(Kovblyuk et al. 2009)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2009)
speciesimage
male Palpal tibia, retrolateral view
(Kovblyuk et al. 2009)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2009)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2009)

world

add

cp

19. Haplodrassus parvicorpus (Roewer, 1951)

speciesimage
male Pedipalp, ventral view
(Bosmans & Van Keer 2012)
speciesimage
male Pedipalp, ventral view
(Bosmans & Van Keer 2012)
speciesimage
male Apical part of bulbus
(Denis 1956)
speciesimage
male Palpal tibia, dorsal view
(Denis 1956)
speciesimage
female Epigyne, ventral view
(Denis 1956)

world

add

cp

20. Haplodrassus ponomarevi Kovblyuk & Seyyar, 2009

speciesimage
male Pedipalp, retrolateral view
(Kovblyuk et al. 2009)
speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2009)
speciesimage
male Palpal tibia, retrolateral view
(Kovblyuk et al. 2009)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2009)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2009)

world

add

cp

21. Haplodrassus pseudosignifer Marusik, Hippa & Koponen, 1996

speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2012b)
speciesimage
male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage
male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2012b)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2012b)

world

add

cp

22. Haplodrassus pugnans (Simon, 1880)

speciesimage
male Pedipalp, ventral view
(Kovblyuk et al. 2012b)
speciesimage
male Bulbus, apical view
(Kovblyuk et al. 2012b)
speciesimage
male Tibial apophysis, retrolateral view
(Kovblyuk et al. 2012b)
speciesimage
female Epigyne, ventral view
(Kovblyuk et al. 2012b)
speciesimage
female Vulva, dorsal view
(Kovblyuk et al. 2012b)

world

add

cp

23. Haplodrassus severus (C. L. Koch, 1839)

speciesimage
male Pedipalp, ventral view
(Simon 1914)
speciesimage
female Epigyne
(Simon 1914)
speciesimage
female Epigyne, variation
(Simon 1914)
24. Haplodrassus umbratilis gothicus Lohmander, 1942 world