Gnaphosidae: Micaria

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1

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Femur I dark brown, distal half light yellow with black-brown spot, a bristle within (often broken off); male pedipalp with narrow cymbium, ventrally with 2 and basally with one bristle; retinaculum narrow; epigyne with a bent ridge; receptaculum seminis oval, ducts divergent; pedipalp femur very large

Micaria dives (Lucas, 1846)

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male Pedipalp
(Thaler 1981c)
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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp
(Bosmans & Blick 2000)
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male Pedipalp
(Thaler 1981c)
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male Bulbus, ventral view
(Thaler 1981c)
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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Braendegaard 1966)
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female Epigyne
(Levy 1973)
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female Vulva, ventral view
(Thaler 1981c)
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female Vulva
(Bosmans & Blick 2000)
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female Vulva, ventral view
(Thaler 1981c)
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female Vulva
(Levy 1973)
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Habitus
(Braendegaard 1966)
- Femur I without black-brown spot on bright background; sexual organ different (cymbium only in subopaca with 2 distal bristles

  2  

2
(1)

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Chelicerae with long grey hairs and with conspicuously, mostly dense copper-coloured scale hairs

Micaria fulgens (Walckenaer, 1802)

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male Pedipalp
(Reimoser 1937)
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male Pedipalp
(Roberts 1995)
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male Pedipalp
(Roberts 1995)
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male Cymbium
(Reimoser 1937)
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female Epigyne
(Braendegaard 1966)
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female Epigyne
(Miller 1971)
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female Vulva
(Miller 1971)
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female Vulva
(Reimoser 1937)
- Chelicerae only with grey, bristle-like hairs

  3  

3
(2)

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Femur I and II distally with 2 bristles; male pedipalp with 1 or 2 small hump-shaped tibial apophysis; cymbium with 3 bristles; embolus very thick; retinaculum missing; epigyne without transverse ridge, medially with variable longitudinal bar

Micaria rossica Thorell, 1875

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male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1988)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp
(Pesarini 2000)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne
(Pesarini 2000)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)
- Femur I and II mostly with one dorsal bristle (only in aenea with 2 bristles); male pedipalp with 1 or 2 larger tibial apophyses; epigyne different

  4  

4
(3)

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Tibial apophysis of male pedipalp dorsally bent; cymbium moderately long, distally with 4 bristles; embolus screw-like; retinaculum large; epigyne with glasses-like structure

Micaria guttulata (C. L. Koch, 1839)

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male Pedipalp, ventral view
(Miller 1967)
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male Pedipalp
(Miller 1967)
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male Pedipalp
(Roberts 1998)
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male Pedipalp
(Reimoser 1937)
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male Cymbium
(Reimoser 1937)
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female Epigyne
(Roberts 1998)
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female Epigyne
(Roberts 1998)
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female Vulva
(Miller 1971)
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female Vulva
(Reimoser 1937)
- Different

  5  

5
(4)

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Femur I and II distally with 2 bristles; male pedipalp with reduced retinaculum; cymbium with 4 bristles; epigyne with large, anteriorly very broad groove

Micaria aenea Thorell, 1871

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male Pedipalp
(Miller 1967)
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male Pedipalp, ventral view
(Miller 1967)
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male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1988)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Vulva
(Miller 1967)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Vulva, ventral view
(Tullgren 1946)
- Femur I and II with one dorsal bristle; retinaculum not reduced; epigyne different

  6  

6
(5)
Cymbium distally with 3 bristles; epigyne with 2 shallow grooves, arcuately bordered or with one large groove and strongly sclerotised margin

  7  

- Cymbium distally with 4 bristles; epigyne without or with small groove

  9  

7
(6)

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Femur III and IV with one distal bristle in basal half; male pedipalp as fig.; epigyne with large, deep groove, its anterior margin strongly sclerotised

Micaria alpina L. Koch, 1872

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male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1988)
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male Pedipalp, ventral view
(Locket & Millidge 1951)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp
(Locket & Millidge 1951)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne
(Locket & Millidge 1951)
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female Epigyne
(Locket et al. 1974)
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female Epigyne
(Locket et al. 1974)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)
- Femur III and IV with 2 distal bristles; sexual organ different

  8  

8
(7)

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Male pedipalp with long and narrow tibial apophysis; embolus thick; distal retinaculum larger; epigyne with shallow groove and V-shaped structure (transverse ridges) in the centre

Micaria lenzi Bösenberg, 1899

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male Pedipalp
(Roberts 1998)
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male Pedipalp
(Reimoser 1937)
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male Cymbium
(Reimoser 1937)
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female Epigyne
(Roberts 1998)
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female Vulva
(Reimoser 1937)
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Male pedipalp with shorter tibial apophysis; embolus thin; retinaculum small, in the middle of bulbus; epigyne as fig.

Micaria silesiaca L. Koch, 1875

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male Pedipalp
(Locket et al. 1974)
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male Pedipalp, ventral view
(Miller 1971)
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male Pedipalp, ventral view
(Locket et al. 1974)
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male Pedipalp
(Reimoser 1937)
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female Epigyne
(Miller 1971)
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female Epigyne/vulva
(Thaler & Noflatscher 1990)
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female Epigyne
(Locket & Millidge 1953)
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female Vulva
(Miller 1967)
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female Vulva
(Reimoser 1937)
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female Vulva, dorsal view
(Tullgren 1946)
9
(6)
Male pedipalp with 2 ± towering tibial apophysis; female tibia I and II ventrally with 1 or 2 bristles; sclerotised epigynal ridge very broad

  10  

- Male pedipalp with 1 tibial apophysis, appressed on cymbium; female tibia I and II ventrally without (pulicaria) or with 3 or 5 pairs of bristles respectively; sclerotised epigynal ridge narrow

  11  

10
(9)

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Prosoma mostly distinctly red-brown (remove scale hairs), very elongated, 1.75-2.0 times as long as wide; opisthosoma ventrally without white longitudinal band, often with white dots; male pedipalp as fig.; position of cymbial bristles variable; epigyne with longish oval receptaculum seminis, reaching to transverse ridge at least

Micaria formicaria (Sundevall, 1831)

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male Pedipalp
(Azheganova 1968)
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male Pedipalp
(Pesarini unpubl.)
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male Pedipalp
(Roberts 1995)
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male Cymbium
(Reimoser 1937)
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female Epigyne
(Miller 1971)
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female Epigyne
(Pesarini unpubl.)
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female Vulva
(Azheganova 1968)
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female Vulva
(Miller 1967)
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female Vulva
(Reimoser 1937)
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female Vulva, dorsal view
(Tullgren 1946)
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female Vulva, ventral view
(Tullgren 1946)
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Prosoma mostly black-brown (remove scale hairs), only 1.3-1.8 times as long as wide; opisthosoma ventrally with white longitudinal band (can be abraded); cymbial bristles variable; epigyne with egg-shaped receptaculum seminis, not reaching to transverse ridge

Micaria coarctata (Lucas, 1846)

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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp, ventral view
(Chyzer & Kulczyński 1897)
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male Pedipalp
(Levy 2002)
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male Pedipalp
(Bosmans & Blick 2000)
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male Pedipalp
(Chyzer & Kulczyński 1897)
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male Pedipalp
(Levy 2002)
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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Chyzer & Kulczyński 1897)
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female Epigyne
(Levy 2002)
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female Epigyne
(Simon 1932)
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female Vulva
(Bosmans & Blick 2000)
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female Vulva, dorsal view
(Tullgren 1946)
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female Spermatheca
(Levy 2002)
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Prosoma, dorsal view
(Levy 2002)
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Eyes, dorsal view
(Levy 2002)
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Opisthosoma, dorsal view
(Levy 2002)
11
(9)

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Male pedipalp with very elongated retinaculum; pedipalp tibia dorsally sometimes with hump; female tibia I and II ventrally without bristles

Micaria pulicaria (Sundevall, 1831)

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male Pedipalp
(Braendegaard 1966)
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male Pedipalp, ventral view
(Braendegaard 1966)
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male Pedipalp (courtesy of the AMNH), ventral view
(Platnick & Shadab 1988)
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male Pedipalp, ventral view
(Locket & Millidge 1951)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp
(Locket & Millidge 1951)
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female Epigyne
(Braendegaard 1966)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne
(Locket & Millidge 1951)
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female Vulva
(Miller 1967)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Vulva, ventral view
(Tullgren 1946)
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female Habitus
(Tullgren 1946)
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Eyes, dorsal view
(Locket & Millidge 1951)
- Male pedipalp with distinctly shorter retinaculum; female tibia I and II ventrally with 3 or 5-6 pairs of bristles

  12  

12
(11)

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Male pedipalp with very strongly bent embolus, without excrescence between embolus and retinaculum; female tibia I and II ventrally with 5-6 pairs of thin bristles; epigyne variable, with bent, mostly bipartite transverse ridge, without groove, posteriorly with sickle-shaped structure

Micaria nivosa L. Koch, 1866

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male Pedipalp
(Mikhailov 1987)
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male Pedipalp, ventral view
(Aakra unpubl.)
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male Pedipalp, ventral view
(Miller 1967)
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male Pedipalp, ventral view
(Mikhailov 1987)
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male Pedipalp, lateral view
(Aakra unpubl.)
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male Pedipalp
(Miller 1967)
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male Pedipalp
(Mikhailov 1987)
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female Epigyne
(Miller 1971)
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female Vulva
(Miller 1967)
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female Vulva, dorsal view
(Tullgren 1946)
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Embolus and retinaculum less bent, with a excrescence between embolus and retinaculum; female tibia I and II ventrally with 3 pairs of robust bristles; epigyne with a transverse ridge within a groove, posteriorly without sickle-shaped structure

Micaria albovittata (Lucas, 1846)

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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp, ventral view
(Levy 2002)
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male Pedipalp, ventral view
(Locket & Millidge 1951)
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male Pedipalp
(Bosmans & Blick 2000)
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male Pedipalp, retrolateral view
(Levy 2002)
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male Pedipalp
(Locket & Millidge 1951)
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male Pedipalp
(Roberts 1995)
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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Levy 2002)
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female Epigyne
(Roberts 1995)
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female Epigyne
(Locket & Millidge 1951)
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female Vulva
(Bosmans & Blick 2000)
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female Vulva
(Miller 1967)
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female Spermathecae, dorsal view
(Levy 2002)
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Prosoma, dorsal view
(Levy 2002)
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Eyes, dorsal view
(Levy 2002)
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Eyes, dorsal view
(Locket & Millidge 1951)
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Metatarsus and tarsus
(Locket & Millidge 1951)
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Opisthosoma, dorsal view
(Levy 2002)
Further taxa

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1. Micaria blicki Kovblyuk & Nadolny, 2008

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male Pedipalp, retrolateral view
(Kovblyuk & Nadolny 2008)
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male Pedipalp, ventral view
(Kovblyuk & Nadolny 2008)
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male Bulbus, ventral view
(Kovblyuk & Nadolny 2008)
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male Median apophysis
(Kovblyuk & Nadolny 2008)
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male Palpal tibia, retrolateral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, dorsal view
(Kovblyuk & Nadolny 2008)

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2. Micaria bosmansi Kovblyuk & Nadolny, 2008

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male Pedipalp, retrolateral view
(Kovblyuk & Nadolny 2008)
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male Pedipalp, ventral view
(Kovblyuk & Nadolny 2008)
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male Bulbus, ventral view
(Kovblyuk & Nadolny 2008)
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male Median apophysis
(Kovblyuk & Nadolny 2008)
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male Palpal tibia, retrolateral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)
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female Epigyne, ventral view
(Kovblyuk & Nadolny 2008)
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female Vulva, ventral view
(Kovblyuk & Nadolny 2008)
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female Vulva, dorsal view
(Kovblyuk & Nadolny 2008)

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3. Micaria constricta Emerton, 1894

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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp, retrolateral view
(Holm 1958b)
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male Pedipalp, ventral view
(Holm 1958b)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne, ventral view
(Holm 1958b)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Vulva
(Holm 1958b)

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4. Micaria donensis Ponomarev & Tsvetkov, 2006

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male Pedipalp, ventral view
(Ponomarev & Tsvetkov 2006)
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male Pedipalp, lateral view
(Ponomarev & Tsvetkov 2006)
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female Epigyne, ventral view
(Ponomarev & Tsvetkov 2006)

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5. Micaria funerea Simon, 1878

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male Pedipalp
(Simon 1932)
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female Epigyne
(Simon 1932)

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6. Micaria gulliae Tuneva & Esyunin, 2003

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male Pedipalp, dorsal view
(Tuneva & Esyunin 2003a)
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male Pedipalp, ventral view
(Tuneva & Esyunin 2003a)
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male Pedipalp, lateral view
(Tuneva & Esyunin 2003a)
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female Epigyne
(Tuneva & Esyunin 2003a)
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female Vulva
(Tuneva & Esyunin 2003a)

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7. Micaria guttigera Simon, 1878

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male Pedipalp
(Simon 1932)
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female Epigyne
(Simon 1932)

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8. Micaria ignea O. P.-Cambridge, 1872

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male Pedipalp, ventral view
(Chatzaki et al. 2002a)
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male Pedipalp, retrolateral view
(Chatzaki et al. 2002a)
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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp, retrolateral view
(Bosmans & Blick 2000)
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female Epigyne, ventral view
(Chatzaki et al. 2002a)
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female Epigyne, ventral view
(Bosmans & Blick 2000)
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female Vulva, dorsal view
(Chatzaki et al. 2002a)
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female Vulva
(Bosmans & Blick 2000)

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9. Micaria kopetdaghensis Mikhailov, 1986

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male Pedipalp, ventral view
(Mikhailov 1987)
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male Pedipalp, ventral view
(Mikhailov 1987)
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male Pedipalp, retrolateral view
(Mikhailov 1987)
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male Pedipalp, prolateral view
(Mikhailov 1987)
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male Palpal tibia, dorsal view
(Mikhailov 1987)
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female Epigyne, ventral view
(Mikhailov 1987)
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female Epigyne, ventral view
(Mikhailov 1987)

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10. Micaria pallipes (Lucas, 1846)

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male Pedipalp, ventral view
(Bosmans 1999)
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male Pedipalp, ventral view
(Levy 2002)
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male Pedipalp, ventral view
(Mikhailov 1987)
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male Pedipalp
(Bosmans & Blick 2000)
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male Pedipalp, retrolateral view
(Levy 2002)
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male Pedipalp
(Mikhailov 1987)
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male Tibial apophysis
(Mikhailov 1987)
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female Epigyne
(Bosmans & Blick 2000)
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female Epigyne
(Levy 2002)
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female Vulva
(Bosmans & Blick 2000)
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female Spermathecae, dorsal view
(Levy 2002)
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Opisthosoma, dorsal view
(Levy 2002)

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11. Micaria palmgreni Wunderlich, 1980

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female Pedipalp, retrolateral view
(Wunderlich 1980g)
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female Epigyne, ventral view
(Wunderlich 1980g)
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female Vulva, dorsal view
(Wunderlich 1980g)

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12. Micaria triguttata Simon, 1884

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male Pedipalp, ventral view
(Bosmans & Blick 2000)
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male Pedipalp
(Bosmans & Blick 2000)
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female Epigyne
(Bosmans 1999)
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female Opisthosoma, dorsal view
(Ledoux et al. 2003)

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13. Micaria tripunctata Holm, 1978

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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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male Pedipalp (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Epigyne (courtesy of the AMNH)
(Platnick & Shadab 1988)
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female Vulva (courtesy of the AMNH)
(Platnick & Shadab 1988)