Thomisidae: Ozyptila

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1
Males

  2  

- Females

  15  

2
(1)
Pedipalp with a ± pointed tibial apophysis, at least 1/3 of cymbium length

  4  

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
- Pedipalp with a ± triangular, blunt tibial apophysis

  3  

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
3
(2)

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cp

Lateral tibial apophysis transverse to longitudinal axis of pedipalp; median tibial apophysis short and broad

Ozyptila scabricula (Westring, 1851)

female

speciesimage
male Pedipalp, ventral view
(Azheganova 1968)
speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Habitus
(Azheganova 1968)
-

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cp

Lateral tibial apophysis diagonal to longitudinal axis of pedipalp; median tibial apophysis narrow and long

Ozyptila sanctuaria (O. P.-Cambridge, 1871)

female

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
4
(2)

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cp

Lateral tibial apophysis transverse to longitudinal axis of pedipalp

Ozyptila brevipes (Hahn, 1826)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
- Different

  5  

5
(4)

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cp

Lateral tibial apophysis tripartite

Ozyptila atomaria (Panzer, 1801)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
- Different

  6  

6
(5)
Lateral tibial apophysis short; median a transverse, S-shaped apophysis

  7  

speciesimage
male Pedipalp, ventral view
(Hippa et al. 1986)
speciesimage
male Pedipalp, ventral view
(Bell & Merrett 2000)
- Different

  10  

7
(6)
Ventral main apophysis transversely S-shaped, ventrally round

  9  

- Different

  8  

8
(7)

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Ventral main apophysis less S-shaped than transversely arrow-shaped; tip of embolus as fig.

Ozyptila secreta Thaler, 1987

female

speciesimage
male Pedipalp, ventral view
(Thaler 1987b)
speciesimage
male Pedipalp, ventral view
(Thaler 1987b)
speciesimage
male Pedipalp, ventral view
(Thaler 1987b)
speciesimage
male Tip of embolus, dorsal view
(Thaler 1987b)
-

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Pedipalp and tip of embolus as fig.

Ozyptila ladina Thaler & Zingerle, 1998

female

speciesimage
male Pedipalp
(Thaler & Zingerle 1998)
speciesimage
male Pedipalp, ventral view
(Thaler & Zingerle 1998)
speciesimage
male Tip of embolus, retrolateral view
(Thaler & Zingerle 1998)
speciesimage
male Tip of embolus, retrolateral view
(Thaler & Zingerle 1998)
speciesimage
male Tibial apophysis
(Thaler & Zingerle 1998)
9
(7)

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Tegular apophysis as fig.

Ozyptila rauda Simon, 1875

female

speciesimage
male Pedipalp, ventral view
(Hippa et al. 1986)
speciesimage
male Pedipalp, ventral view
(Roberts 1998)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
speciesimage
male Tegular apophysis, prolateral view
(Hippa et al. 1986)
-

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Tegular apophysis as fig.

Ozyptila pullata (Thorell, 1875)

female

speciesimage
male Pedipalp, ventral view
(Bell & Merrett 2000)
speciesimage
male Pedipalp, ventral view
(Hippa et al. 1986)
speciesimage
male Pedipalp, ventral view
(Pozzi & Hänggi 1998)
speciesimage
male Pedipalp, ventral view
(Roberts 1998)
speciesimage
male Tegular apophysis, prolateral view
(Hippa et al. 1986)
10
(6)
Lateral tibial apophysis angled or outwardly bent

  11  

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
- Lateral tibial apophysis straight or inwardly bent

  12  

11
(10)

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cp

Median tibial apophysis deeply furcate

Ozyptila praticola (C. L. Koch, 1837)

female

speciesimage
male Pedipalp
(Azheganova 1968)
speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
-

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cp

Median tibial apophysis not distinctly furcate

Ozyptila claveata (Walckenaer, 1837)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
12
(10)

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Lateral tibial apophysis diagonal to longitudinal axis of pedipalp

Cozyptila blackwalli (Simon, 1875)

female

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
- Lateral tibial apophysis parallel to longitudinal axis of pedipalp

  13  

13
(12)

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cp

Lateral tibial apophysis approx. 1/2 of cymbium length; median tibial apophysis with 2 processes, pointing to each other in a right angle

Ozyptila trux (Blackwall, 1846)

female

speciesimage
male Pedipalp, ventral view
(Braendegaard 1972a)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp, ventral view
(Wunderlich & Schultz 1995)
speciesimage
male Tegular apophysis
(Wunderlich & Schultz 1995)
-

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Embolus distally a bit stronger bent; cove of distal apophysis larger basal apophysis smaller (cf. Wunderlich 1994)

Ozyptila westringi (Thorell, 1873)

female

speciesimage
male Pedipalp, ventral view
(Roberts 1998)
speciesimage
male Pedipalp, ventral view
(Wunderlich & Schultz 1995)
speciesimage
male Tegular apophysis
(Wunderlich & Schultz 1995)
speciesimage
male Palpal tibia, retrolateral view
(Wunderlich & Schultz 1995)
- Lateral tibial apophysis approx. 3/4 cymbium length; median tibial apophysis with 2 ± parallel processes

  14  

speciesimage
male Pedipalp, ventral view
(Kupryjanowicz 1994b)
speciesimage
male Pedipalp, ventral view
(Roberts 1995)
14
(13)

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cp

Tibial apophysis as fig.

Ozyptila gertschi Kurata, 1944

female

speciesimage
male Pedipalp, ventral view
(Kupryjanowicz 1994b)
speciesimage
male Pedipalp, ventral view
(Roberts 1998)
speciesimage
male Pedipalp
(Kupryjanowicz 1994b)
-

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Tibial apophysis as fig.

Ozyptila simplex (O. P.-Cambridge, 1862)

female

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1951)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
15
(1)
Epigyne with a large, distinctly margined groove

  16  

speciesimage
female Epigyne, ventral view
(Hippa et al. 1986)
speciesimage
female Epigyne, ventral view
(Hippa et al. 1986)
speciesimage
female Epigyne
(Roberts 1995)
- Epigyne without big groove

  20  

16
(15)

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cp

Epigynal groove ± kidney-shaped, largest width near posterior margin

Ozyptila praticola (C. L. Koch, 1837)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Azheganova 1968)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
- Different

  17  

17
(16)

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Lateral borders of epigynal grooves indistinct, long median septum

Ozyptila ladina Thaler & Zingerle, 1998

male

speciesimage
female Epigyne, ventral view
(Thaler & Zingerle 1998)
speciesimage
female Vulva, dorsal view
(Thaler & Zingerle 1998)
- Different

  18  

18
(17)

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Epigynal groove almost completely filled with broad protrusion of anterior margin

Ozyptila secreta Thaler, 1987

male

speciesimage
female Epigyne/vulva, ventral view
(Thaler 1987b)
speciesimage
female Epigyne/vulva, dorsal view
(Thaler 1987b)
- Different

  19  

19
(18)

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Bottom of epigynal groove with transverse folds

Ozyptila rauda Simon, 1875

male

speciesimage
female Epigyne, ventral view
(Hippa et al. 1986)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva, dorsal view
(Hippa et al. 1986)
-

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Bottom of epigynal groove with semicircular folds

Ozyptila pullata (Thorell, 1875)

male

speciesimage
female Epigyne
(Bell & Merrett 2000)
speciesimage
female Epigyne, ventral view
(Hippa et al. 1986)
speciesimage
female Epigyne
(Pozzi & Hänggi 1998)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva, dorsal view
(Hippa et al. 1986)
20
(15)
Epigyne anteriorly with a short, blunt, backwardly directed process

  21  

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Roberts 1995)
- Epigyne anteriorly without such process

  24  

21
(20)

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Epigyne wider than long in anterior epigynal area

Ozyptila trux (Blackwall, 1846)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Wunderlich & Schultz 1995)
-

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Copulatory openings nearer to receptacula (cf. Wunderlich 1994)

Ozyptila westringi (Thorell, 1873)

male

speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne, ventral view
(Wunderlich & Schultz 1995)
speciesimage
female Vulva, dorsal view
(Wunderlich & Schultz 1995)
- Process ± as long as wide

  22  

22
(21)

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cp

Prosoma uniformly dark brown; epigyne as fig.

Ozyptila scabricula (Westring, 1851)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Azheganova 1968)
- Prosoma yellow-brown with 2-4 dark brown longitudinal stripes

  23  

23
(22)

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Distance between posterior median eyes > 2 diameters; body length 4-5 mm; epigyne and vulva as fig.

Ozyptila simplex (O. P.-Cambridge, 1862)

male

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva, ventral view
(Wunderlich 1973b)
-

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Distance between posterior median eyes < 2 diameters; body length about 3.7 mm; epigyne and vulva as fig.

Ozyptila gertschi Kurata, 1944

male

speciesimage
female Epigyne
(Kupryjanowicz 1994b)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Vulva
(Kupryjanowicz 1994b)
speciesimage
Habitus
(Kupryjanowicz 1994b)
24
(20)
Opisthosoma dark grey to dark brown

  25  

- Opisthosoma light yellowish to middle brown

  26  

25
(24)

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Prosoma with light median band; epigyne as fig.

Cozyptila blackwalli (Simon, 1875)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
-

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Prosoma uniformly dark; epigyne and vulva as fig.

Ozyptila claveata (Walckenaer, 1837)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
26
(24)

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Epigyne with slantingly running folds in anterior region

Ozyptila sanctuaria (O. P.-Cambridge, 1871)

male

speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
- Different

  27  

27
(26)

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Epigyne bipartite in anterior region

Ozyptila atomaria (Panzer, 1801)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva, ventral view
(Heimer & Nentwig 1991)
-

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Epigyne not bipartite, in anterior part with a small transverse groove, posterior to it a bulged region

Ozyptila brevipes (Hahn, 1826)

male

speciesimage
female Epigyne
(Braendegaard 1972a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
Further taxa

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1. Ozyptila ankarensis Karol, 1966

speciesimage
male Pedipalp, ventral view
(Karol 1966b)

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2. Ozyptila arctica Kulczyński, 1908

speciesimage
male Pedipalp, ventral view
(Roberts 1995)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne, ventral view
(Løvbrekke unpubl.)
speciesimage
female Vulva, dorsal view
(Løvbrekke unpubl.)
speciesimage
female Prosoma, dorsal view
(Løvbrekke unpubl.)
speciesimage
female Sternum
(Løvbrekke unpubl.)
speciesimage
female Habitus, dorsal view
(Løvbrekke unpubl.)

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3. Ozyptila balcanica Deltshev, Blagoev, Komnenov & Lazarov, 2016

speciesimage
male Pedipalp, prolateral view
(Deltshev et al. 2016)
speciesimage
male Pedipalp, retrolateral view
(Deltshev et al. 2016)
speciesimage
male Pedipalp, ventral view
(Deltshev et al. 2016)
speciesimage
female Epigyne
(Deltshev et al. 2016)
speciesimage
female Vulva
(Deltshev et al. 2016)

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4. Ozyptila bejarana Urones, 1998

speciesimage
male Pedipalp, ventral view
(Urones 1998)
speciesimage
male Pedipalp
(Urones 1998)
speciesimage
male Tibial apophysis
(Urones 1998)
speciesimage
male Habitus
(Urones 1998)
speciesimage
female Epigyne
(Urones 1998)
speciesimage
female Vulva
(Urones 1998)
speciesimage
female Habitus
(Urones 1998)

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5. Ozyptila clavidorsa Roewer, 1959

speciesimage
female Epigyne, ventral view
(Roewer 1959a)
speciesimage
female Leg I, dorsal view
(Roewer 1959a)

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6. Ozyptila confluens (C. L. Koch, 1845)

speciesimage
male Pedipalp, ventral view
(Simon 1932)
speciesimage
male Palpal tibia
(Simon 1932)
speciesimage
female Epigyne
(Simon 1932)
speciesimage
female Epigyne, ventral view
(Kolosváry 1939b)
speciesimage
female Epigyne
(Breitling et al. 2016b)

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7. Ozyptila conostyla Hippa, Koponen & Oksala, 1986

speciesimage
male Pedipalp, ventral view
(Marusik & Logunov 1995a)
speciesimage
male Pedipalp, retrolateral view
(Marusik & Logunov 1995a)
speciesimage
male Tip of embolus, ventral view
(Marusik & Logunov 1995a)
speciesimage
male Tegular apophysis, prolateral view
(Marusik & Logunov 1995a)

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8. Ozyptila dagestana Ponomarev & Dvadnenko, 2011

speciesimage
male Pedipalp, ventral view
(Ponomarev et al. 2011)
speciesimage
male Bulbus, ventral view
(Ponomarev et al. 2011)
speciesimage
female Epigyne, ventral view
(Ponomarev et al. 2011)

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9. Ozyptila danubiana Weiss, 1998

speciesimage
male Pedipalp, ventral view
(Weiss 1998a)
speciesimage
male Pedipalp, retrolateral view
(Weiss 1998a)
speciesimage
male Pedipalp, aboral view
(Weiss 1998a)

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10. Ozyptila elegans (Blackwall, 1870)

speciesimage
male Pedipalp, ventral view
(Blackwall 1870b)
speciesimage
male Habitus, dorsal view
(Blackwall 1870b)

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11. Ozyptila furcula L. Koch, 1882

speciesimage
male Pedipalp, ventral view
(Simon 1932)
speciesimage
male Pedipalp, retrolateral view
(Lecigne 2014)
speciesimage
male Pedipalp, ventral view
(Lecigne 2014)
speciesimage
male Palpal tibia
(Simon 1932)
speciesimage
female Epigyne
(Simon 1932)

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12. Ozyptila lugubris (Kroneberg, 1875)

speciesimage
male Pedipalp, ventral view
(Marusik & Logunov 1990)
speciesimage
male Pedipalp, retrolateral view
(Marusik & Logunov 1990)
speciesimage
male Pedipalp, ventral view
(Marusik & Logunov 1990)
speciesimage
male Palpal tibia, retrolateral view
(Marusik & Logunov 1990)
speciesimage
female Epigyne, ventral view
(Marusik & Logunov 1990)
speciesimage
female Epigyne, variation, ventral view
(Marusik & Logunov 1990)
speciesimage
female Epigyne, variation, ventral view
(Marusik & Logunov 1990)

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13. Ozyptila maculosa Hull, 1948

speciesimage
male Palpal tibia, ventral view
(Hull 1948)

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14. Ozyptila mingrelica Mcheidze, 1971

speciesimage
female Epigyne, ventral view
(Mcheidze 2014)
speciesimage
female Habitus, dorsal view
(Mcheidze 2014)

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15. Ozyptila orientalis balkarica Ovtsharenko, 1979

speciesimage
male Pedipalp, retrolateral view
(Marusik 2008)
speciesimage
male Pedipalp, prolateral view
(Marusik 2008)
speciesimage
male Pedipalp, apical view
(Marusik 2008)
speciesimage
male Pedipalp, retrolateral view
(Ovtsharenko 1979)
speciesimage
male Pedipalp, ventral view
(Ovtsharenko 1979)
speciesimage
male Tegular apophysis, prolateral view
(Marusik 2008)
speciesimage
female Epigyne
(Ovtsharenko 1979)
speciesimage
female Vulva, dorsal view
(Logunov & Marusik 1994b)

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16. Ozyptila orientalis basegica Esyunin, 1992

speciesimage
male Pedipalp, ventral view
(Esyunin 1992a)
speciesimage
male Pedipalp, lateral view
(Esyunin 1992a)
speciesimage
male Tegular apophysis
(Esyunin 1992a)
speciesimage
male Habitus, dorsal view
(Esyunin 1992a)
speciesimage
female Epigyne, ventral view
(Esyunin 1992a)
speciesimage
female Vulva
(Esyunin 1992a)

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17. Ozyptila pauxilla (Simon, 1870)

speciesimage
male Pedipalp, ventral view
(Simon 1932)
speciesimage
male Pedipalp, retrolateral view
(Lecigne 2012)
speciesimage
male Pedipalp, ventral view
(Lecigne 2012)
speciesimage
male Palpal tibia
(Simon 1932)
speciesimage
female Epigyne, ventral view
(Denis 1937b)

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18. Ozyptila perplexa Simon, 1875

speciesimage
male Pedipalp, ventral view
(Simon 1932)
speciesimage
male Palpal tibia
(Simon 1932)

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19. Ozyptila rigida (O. P.-Cambridge, 1872)

speciesimage
male Pedipalp, ventral view
(Levy 1985)
speciesimage
male Pedipalp, retrolateral view
(Levy 1985)
speciesimage
female Epigyne, ventral view
(Kulczyński 1908a)
speciesimage
female Epigyne, ventral view
(Levy 1985)
speciesimage
female Vulva, dorsal view
(Levy 1985)

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20. Ozyptila salustri Wunderlich, 2011

speciesimage
male Pedipalp, ventral view
(Wunderlich 2011a)
speciesimage
male Pedipalp, retrolateral view
(Wunderlich 2011a)
speciesimage
male Palpal tibia, dorsal view
(Wunderlich 2011a)

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21. Ozyptila sincera Kulczyński, 1926

speciesimage
male Pedipalp, ventral view
(Ono 1988)
speciesimage
male Pedipalp, retrolateral view
(Ono 1988)
speciesimage
female Epigyne, ventral view
(Ono 1988)
speciesimage
female Vulva, dorsal view
(Ono 1988)

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22. Ozyptila spirembola Wunderlich, 1995

speciesimage
male Pedipalp, ventral view
(Wunderlich 1995b)
speciesimage
male Pedipalp, retrolateral view
(Wunderlich 1995b)

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23. Ozyptila tricoloripes Strand, 1913

speciesimage
male Pedipalp, ventral view
(Levy 1985)
speciesimage
male Pedipalp, retrolateral view
(Levy 1985)
speciesimage
female Epigyne, ventral view
(Demir et al. 2008b)
speciesimage
female Epigyne, ventral view
(Levy 1985)
speciesimage
female Vulva, dorsal view
(Demir et al. 2008b)
speciesimage
female Vulva, dorsal view
(Levy 1985)

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24. Ozyptila umbraculorum Simon, 1932

speciesimage
male Pedipalp, ventral view
(Jiménez-Valverde 2002)
speciesimage
male Pedipalp, prolateral view
(Deltshev et al. 2016)
speciesimage
male Pedipalp, retrolateral view
(Deltshev et al. 2016)
speciesimage
male Pedipalp, ventral view
(Deltshev et al. 2016)
speciesimage
female Epigyne
(Urones 1985)
speciesimage
female Epigyne
(Deltshev et al. 2016)
speciesimage
female Vulva
(Urones 1985)
speciesimage
female Vulva
(Deltshev et al. 2016)
25. Ozyptila flava Simon, 1875 world
26. Ozyptila nigristerna Dalmas, 1922 world
27. Ozyptila trux devittata Strand, 1901 world