Lycosidae: Arctosa

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1
Males

  2  

- Females

  11  

2
(1)

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Prosoma lateral with 3-4 spots; prosoma length of males 3.1-3.2 mm; prosoma olive-brown, median band bright, weakly star-shaped; legs red-brown; coxa and femur yellowish; opisthosoma yellow with single black hairs; cardiac mark distinct, white haired; posterior to cardiac mark with 4-5 bright, triangular spots, laterally 2 yellow spots; pedipalp as fig.

Arctosa lutetiana (Simon, 1876)

female

speciesimage
male Pedipalp
(Wiehle 1967)
speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
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male Pedipalp, ventral view
(Buchar & Zdárek 1960)
speciesimage
male Pedipalp, ventral view
(Roberts 1998)
speciesimage
male Pedipalp, ventral view
(Wiehle 1967)
speciesimage
male Pedipalp, ventral view
(Loksa 1972)
speciesimage
male Pedipalp, ventral view
(Wiebes 1959)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp
(Roberts 1998)
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Chelicerae, frontal view
(Wiehle 1967)
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Chelicerae, ventral view
(Wiehle 1967)
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Opisthosoma, dorsal view
(Wiehle 1967)
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Habitus
(Lugetti & Tongiorgi 1965)
- Different

  3  

3
(2)

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Cardiac mark on opisthosoma broadened anteriorly; prosoma length 3.6-3.8 mm; prosoma red-brown, with black dots; legs reddish with dark spots on femur; opisthosoma reddish; cardiac mark white-yellowish, bordered with black, posterior to cardiac mark 2-3 dark spots, laterally 2-3 round, black spots from which running fine white lines; male pedipalp as fig.

Arctosa alpigena (Doleschall, 1852)

female

speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp, ventral view
(Locket et al. 1974)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Roberts 1998)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
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Tarsus of leg I
(Buchar 1981)
- Different

  4  

4
(3)

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Pedipalp tibia distally with 2 projections; prosoma length about 3.8 mm; prosoma yellow-reddish, medially with serrated band, laterally with stripes consisting of 4-5 bright, irregular and ± merging dots; legs yellowish, annulated basally

Arctosa figurata (Simon, 1876)

female

speciesimage
male Pedipalp
(Roberts 1998)
speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Roberts 1998)
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Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1965)
- Pedipalp tibia distally without projections

  5  

5
(4)
Tegular apophysis ± triangular due to ventral fold, bent toward median part of pedipalp

  6  

- Tegular apophysis ends with a ± finger-shaped, basally directed process

  9  

6
(5)
Spines on metatarsus I and II conspicuously short; spines on tibia I reduced or lacking; cheliceral furrow margin with 2 or 3 teeth on inner side; upper part of bulbus with 2 ± pointed projections

  7  

- Spines on metatarsus I-II and tibia I not conspicuously reduced or lacking; cheliceral furrow margin with 3 teeth on inner side; upper part of bulbus with 1 ± pointed projection

  8  

7
(6)

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Cheliceral furrow margin with 2 teeth; pedipalp as fig.; prosoma length 5.3-6.0 mm; prosoma red-brown, light median band weakly serrated, lateral bands narrow, indistinct and often broken; legs reddish yellow; femur spotted with brown

Arctosa fulvolineata (Lucas, 1846)

female

speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
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male Pedipalp, lateral view
(Lugetti & Tongiorgi 1965)
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male Pedipalp
(Roberts 1998)
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male Pedipalp
(Locket & Millidge 1951)
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female Habitus
(Lugetti & Tongiorgi 1965)
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Cheliceral furrow margin with 3 teeth; pedipalp as fig; prosoma length 3.8-4.8 mm; prosoma reddish, bright median band broad anteriorly, narrow and serrated posteriorly, lateral bands striking; legs yellow-reddish, annulated

Arctosa personata (L. Koch, 1872)

female

speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
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male Pedipalp
(Heimer & Nentwig 1991)
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Habitus
(Heimer & Nentwig 1991)
8
(6)

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Prosoma compact, plump; prosoma length 4.7-5.4 mm; basic colouration greenish yellow; legs indistinctly annulated; pedipalp as fig.

Arctosa cinerea (Fabricius, 1777)

female

speciesimage
male Pedipalp, ventral view
(Aakra unpubl.)
speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
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male Pedipalp
(Azheganova 1968)
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male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
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male Pedipalp
(Roberts 1998)
speciesimage
male Pedipalp
(Loksa 1972)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
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male Tegular apophysis
(Loksa 1972)
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Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1965)
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Prosoma narrow, slender; prosoma length 3.0-3.9 mm; colouration brown; legs distinctly annulated; pedipalp as fig.

Arctosa perita (Latreille, 1799)

female

speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp, ventral view
(Wiebes 1959)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Roberts 1998)
speciesimage
male Pedipalp
(Loksa 1972)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Tegular apophysis
(Loksa 1972)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1969)
9
(5)

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Yellowish median stripe on opisthosoma parallel-sided in anterior part, distinctly marked off from remaining dark opisthosomal colour; tegular apophysis medially with triangular and pointed projection; prosoma length 3.0-4.1 mm

Arctosa leopardus (Sundevall, 1833)

female

speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp, ventral view
(Wiebes 1959)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Roberts 1998)
speciesimage
male Pedipalp
(Loksa 1972)
speciesimage
male Pedipalp
(Locket & Millidge 1951)
speciesimage
male Embolus
(Loksa 1972)
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Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1965)
- Median stripe on opisthosoma with broadened anterior part

  10  

10
(9)

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Tegular apophysis with process, pointing basally; prosoma length 2.7-3.7 mm; prosoma yellow-brown, whitish in eye region, medially with light serrated band, lateral stripes striking, consisting of ± merging single dots; legs yellowish, indistinctly annulated

Arctosa stigmosa (Thorell, 1875)

female

speciesimage
male Pedipalp, ventral view
(Aakra 2000d)
speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Heimer & Nentwig 1991)
speciesimage
male Habitus
(Aakra 2000d)
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Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Heimer & Nentwig 1991)
-

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Tegular apophysis with a process, pointing distally; prosoma length 4.1-5.0 mm; prosoma red-brown with dark spots, brighter median band distinctly serrated, 2 bright spots posterior to posterior lateral eyes, lateral stripes consisting of bright spots; legs yellowish; femur weakly annulated

Arctosa maculata (Hahn, 1822)

female

speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1969)
speciesimage
male Pedipalp
(Loksa 1972)
speciesimage
male Embolus
(Loksa 1972)
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Habitus
(Breuss 2001)
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Habitus
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Lugetti & Tongiorgi 1965)
11
(1)

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Prosoma laterally with 3-4 spots; prosoma length of females 3.4-3.7 mm; prosoma olive brown, median band bright, weakly star-shaped; legs red-brown, coxa and femur yellowish; opisthosoma yellow greenish with scarce black hairs; cardiac mark distinct, white haired, posterior to cardiac mark 4-5 bright triangular spots, lateral 2 yellow spots; epigyne and vulva as fig.

Arctosa lutetiana (Simon, 1876)

male

speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Epigyne, ventral view
(Buchar & Zdárek 1960)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Wiehle 1967)
speciesimage
female Epigyne
(Loksa 1972)
speciesimage
female Epigyne
(Wiebes 1959)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
speciesimage
female Vulva
(Wiehle 1967)
speciesimage
female Vulva
(Loksa 1972)
speciesimage
Habitus
(Heimer & Nentwig 1991)
speciesimage
Habitus
(Lugetti & Tongiorgi 1965)
- Different

  12  

12
(11)

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Cardiac mark on opisthosoma broadened anteriorly; prosoma length 3.8-4.1 mm; prosoma red-brown, with black dots; legs reddish with dark spots on femur; opisthosoma reddish; cardiac mark white-yellowish, bordered with black, posterior to cardiac mark 2-3 dark spots, laterally 2-3 round, black spots from which running fine white lines; epigyne and vulva as figs.

Arctosa alpigena (Doleschall, 1852)

male

speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Lugetti & Tongiorgi 1965)
- Different

  13  

13
(12)

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Median part of epigyne almost as large as epigynal groove

Arctosa figurata (Simon, 1876)

male

speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1965)
- Median part of epigyne small; between the lateral epigynal parts a large epigynal groove visible

  14  

14
(13)

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Cheliceral furrow margin with 2 teeth on inner side

Arctosa fulvolineata (Lucas, 1846)

male

speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Habitus
(Lugetti & Tongiorgi 1965)
- Cheliceral furrow margin with 3 teeth on inner side

  15  

15
(14)

Anterior epigynal pockets closely together; median bar indistinct, triangular, haired

  16  

-

Anterior epigynal pockets distinctly separated; median bar distinctly visible

  18  

16
(15)

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Epigynal groove wider anteriorly than posteriorly; margins of median bar merging with margin of anterior epigynal pocket. Spines on metatarsus I-II short; spines on tibia I reduced or missing

Arctosa personata (L. Koch, 1872)

male

speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
speciesimage
Habitus
(Heimer & Nentwig 1991)
- Epigynal groove posteriorly at least as wide as anteriorly; margins of median bar distinctly separated from margin of epigynal pocket; spines on metatarsus I-II and tarsus normal

  17  

17
(16)

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Prosoma slender and long; basic colouration brown; prosoma length smaller or equal 4.0 mm; anterior margin of epigynal pocket broad and strongly sclerotised

Arctosa perita (Latreille, 1799)

male

speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Loksa 1972)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Epigyne
(Wiebes 1959)
speciesimage
female Vulva
(Loksa 1972)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Lugetti & Tongiorgi 1969)
-

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Prosoma plump; basic colouration greenish yellow. Prosoma length > 4.5 mm; anterior margin of epigynal pocket slender

Arctosa cinerea (Fabricius, 1777)

male

speciesimage
female Epigyne
(Aakra unpubl.)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Loksa 1972)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Vulva
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Vulva
(Loksa 1972)
speciesimage
female Vulva
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Habitus
(Aakra 2000d)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Lugetti & Tongiorgi 1965)
18
(15)

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Median part of epigyne reaching epigastric furrow, triangular

Arctosa leopardus (Sundevall, 1833)

male

speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Roberts 1998)
speciesimage
female Epigyne
(Loksa 1972)
speciesimage
female Epigyne
(Locket & Millidge 1951)
speciesimage
female Epigyne
(Wiebes 1959)
speciesimage
female Vulva
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Vulva
(Loksa 1972)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
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Habitus
(Lugetti & Tongiorgi 1965)
- Median part of epigyne not reaching epigastric furrow

  19  

19
(18)

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Median part of epigyne posteriorly divergent

Arctosa maculata (Hahn, 1822)

male

speciesimage
female Epigyne
(Breuss 2001)
speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Epigyne
(Loksa 1972)
speciesimage
female Vulva
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Vulva
(Loksa 1972)
speciesimage
Habitus
(Breuss 2001)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Lugetti & Tongiorgi 1965)
-

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Median part of epigyne anteriorly divergent

Arctosa stigmosa (Thorell, 1875)

male

speciesimage
female Epigyne
(Aakra 2000d)
speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Vulva
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Heimer & Nentwig 1991)
Further taxa

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1. Arctosa alpigena lamperti Dahl, 1908

speciesimage
male Pedipalp
(Buchar 1981)
speciesimage
male Pedipalp, ventral view
(Buchar 1981)
speciesimage
female Epigyne, ventral view
(Miller 1971)
speciesimage
Tarsus of leg I
(Buchar 1981)

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2. Arctosa intricaria (C. L. Koch, 1847)

speciesimage
male Lamella characteristica
(Roewer 1960b)
speciesimage
male Habitus, dorsal view
(Roewer 1960b)
speciesimage
female Epigyne, ventral view
(Roewer 1960b)
speciesimage
female Eyes, frontal view
(C. L. Koch 1847)
speciesimage
female Habitus
(C. L. Koch 1847)

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3. Arctosa lacustris (Simon, 1876)

speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1969)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp, ventral view
(Simon 1937)
speciesimage
male Pedipalp, ventral view
(Hepner & Paulus 2009)
speciesimage
female Epigyne, ventral view
(Simon 1937)
speciesimage
female Epigyne, ventral view
(Hepner & Paulus 2009)
speciesimage
female Epigyne, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
female Vulva, dorsal view
(Hepner & Paulus 2009)
speciesimage
female Habitus
(Lugetti & Tongiorgi 1965)

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4. Arctosa lagodechiensis Mcheidze, 1997

speciesimage
female Epigyne/vulva
(Mcheidze 1997)
speciesimage
female Habitus, dorsal view
(Mcheidze 1997)

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5. Arctosa letourneuxi (Simon, 1885)

speciesimage
male Pedipalp, ventral view
(Guy 1966)
speciesimage
female Epigyne, ventral view
(Guy 1966)

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6. Arctosa ravida Ponomarev, 2007

speciesimage
female Epigyne, ventral view
(Ponomarev 2007a)

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7. Arctosa renidescens Buchar & Thaler, 1995

speciesimage
male Pedipalp, ventral view
(Buchar 1981)
speciesimage
male Pedipalp
(Buchar 1981)
speciesimage
male Pedipalp, ventral view
(Lugetti & Tongiorgi 1965)
speciesimage
male Pedipalp
(Lugetti & Tongiorgi 1965)
speciesimage
male Bulbus, ventral view
(Buchar & Thaler 1995a)
speciesimage
male Embolic division
(Buchar & Thaler 1995a)
speciesimage
male Prosoma
(Buchar 1981)
speciesimage
female Epigyne
(Buchar 1981)
speciesimage
female Epigyne
(Denis 1963a)
speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Epigyne
(Lugetti & Tongiorgi 1965)
speciesimage
female Vulva
(Buchar 1981)
speciesimage
female Vulva
(Lugetti & Tongiorgi 1965)
speciesimage
Habitus
(Heimer & Nentwig 1991)

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8. Arctosa similis Schenkel, 1938

speciesimage
male Pedipalp, ventral view
(Simon 1937)
speciesimage
male Pedipalp, ventral view
(Hepner & Paulus 2009)
speciesimage
female Epigyne, ventral view
(Simon 1937)
speciesimage
female Epigyne, ventral view
(Hepner & Paulus 2009)
speciesimage
female Vulva, dorsal view
(Hepner & Paulus 2009)

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9. Arctosa simoni Guy, 1966

speciesimage
female Epigyne, ventral view
(Guy 1966)
speciesimage
female Habitus, dorsal view
(Guy 1966)

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10. Arctosa tbilisiensis Mcheidze, 1946

speciesimage
male Pedipalp, ventral view
(Thaler et al. 2000)
speciesimage
male Pedipalp, ventral view
(Alderweireldt 2002)
speciesimage
male Pedipalp, retrolateral view
(Alderweireldt 2002)
speciesimage
male Embolic division, ventral view
(Thaler et al. 2000)
speciesimage
male Embolic division, dorsal view
(Thaler et al. 2000)
speciesimage
male Habitus
(Alderweireldt 2002)
speciesimage
female Epigyne, ventral view
(Alderweireldt 2002)
speciesimage
female Vulva, ventral view
(Alderweireldt 2002)

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11. Arctosa variana C. L. Koch, 1847

speciesimage
male Pedipalp, ventral view
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
male Pedipalp
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
female Epigyne
(Fuhn & Niculescu-Burlacu 1971)
speciesimage
Habitus
(Fuhn & Niculescu-Burlacu 1971)

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12. Arctosa villica (Lucas, 1846)

speciesimage
male Pedipalp, ventral view
(Murphy & Tongiorgi 1979)
speciesimage
male Pedipalp
(Murphy & Tongiorgi 1979)
speciesimage
female Epigyne
(Murphy & Tongiorgi 1979)
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Habitus
(Murphy & Tongiorgi 1979)
13. Arctosa excellens (Simon, 1876) world
14. Arctosa perita arenicola (Simon, 1937) world