Agelenidae: Coelotes

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1
Males

  2  

- Females

  7  

2
(1)
Conductor of pedipalp distally long, reaching beyond cymbium laterally

  3  

- Conductor shorter, not reaching beyond cymbium laterally

  4  

3
(2)

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Tip of conductor transverse to longitudinal axis of cymbium; conductor basally with a broad process

Inermocoelotes inermis (L. Koch, 1855)

female

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
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male Pedipalp
(Dahl 1931)
speciesimage
male Pedipalp
(De Blauwe 1973)
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male Pedipalp
(Roberts 1995)
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male Palpal tibia
(Loksa 1969)
-

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Conductor directed toward tip of cymbium; patella with 2 apophyses

Urocoras longispinus (Kulczyński, 1897)

female

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp, ventral view
(Heimer & Nentwig 1991)
speciesimage
male Pedipalp
(De Blauwe 1973)
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male Pedipalp
(Loksa 1969)
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male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
4
(2)
Conductor distally with a small, stepped tip; patellar apophysis with a short process

  5  

-

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Conductor without stepped tip; patellar apophysis with 3 short, rounded processes; opisthosoma mostly lighter than in terrestris and patter less distinct

Coelotes atropos (Walckenaer, 1830)

female

speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1953)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Pedipalp
(Dahl 1931)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patella and tibia of pedipalp
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(Loksa 1969)
speciesimage
male Patella and tibia of pedipalp
(Locket & Millidge 1953)
5
(4)

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Tip of conductor broad, rounded; distal edge of patellar apophysis straight

Coelotes solitarius L. Koch, 1868

female

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Pedipalp
(Dahl 1931)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(Loksa 1969)
speciesimage
male Patellar apophysis
(Loksa 1969)
- Different

  6  

6
(5)

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Tip of conductor narrow and short; distal edge of patellar apophysis distinctly indented

Coelotes terrestris (Wider, 1834)

female

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp, ventral view
(Locket & Millidge 1953)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Pedipalp
(Roberts 1995)
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male Tip of pedipalp
(Locket & Millidge 1953)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(Loksa 1969)
speciesimage
male Patellar apophysis
(Loksa 1969)
speciesimage
male Patella and tibia of pedipalp
(Locket & Millidge 1953)
-

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Tip of conductor longer, step-like

Coelotes mediocris Kulczyński, 1887

female

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp
(De Blauwe 1973)
7
(1)
Epigyne as long as wide

  8  

- Epigyne a bit longer than wide, ± square

  10  

8
(7)

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Epigyne with parallel lateral margins, anterior margin distinctly marked

Coelotes terrestris (Wider, 1834)

male

speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(Roberts 1995)
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female Epigyne
(Loksa 1969)
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female Epigyne
(Locket & Millidge 1953)
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female Epigyne
(Locket et al. 1974)
speciesimage
female Vulva
(Heimer & Nentwig 1991)
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female Vulva
(Loksa 1969)
- Different

  9  

9
(8)

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Epigyne anteriorly bulged laterally, indistinct; opisthosoma mostly lighter than in terrestris and patter less distinct

Coelotes atropos (Walckenaer, 1830)

male

speciesimage
female Epigyne
(Dahl 1931)
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female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(Loksa 1969)
speciesimage
female Epigyne
(Locket & Millidge 1953)
speciesimage
female Epigyne
(Locket et al. 1974)
speciesimage
female Vulva
(Dahl 1931)
speciesimage
female Vulva
(Loksa 1969)
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Epigyne as fig.

Coelotes mediocris Kulczyński, 1887

male

speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(De Blauwe 1973)
10
(7)

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Lateral margins of epigyne convex, bent outwardly

Urocoras longispinus (Kulczyński, 1897)

male

speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(Heimer & Nentwig 1991)
speciesimage
female Epigyne
(Loksa 1969)
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female Vulva
(Loksa 1969)
- Lateral margins of epigyne concave, bent inwardly

  11  

11
(10)

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Anterior margin of epigyne distinctly marked, rounded

Inermocoelotes inermis (L. Koch, 1855)

male

speciesimage
female Epigynal plate
(Dahl 1931)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne, ventral view
(Jäger 1995a)
speciesimage
female Epigyne, ventral view
(Jäger 1995a)
speciesimage
female Epigyne
(Roberts 1995)
speciesimage
female Epigyne
(Loksa 1969)
speciesimage
female Vulva
(Dahl 1931)
speciesimage
female Vulva
(Loksa 1969)
speciesimage
female Vulva, dorsal view
(Jäger 1995a)
-

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Anterior margin of epigyne indistinct, straight

Coelotes solitarius L. Koch, 1868

male

speciesimage
female Epigyne
(Dahl 1931)
speciesimage
female Epigyne
(Dahl 1931)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(Loksa 1969)
speciesimage
female Vulva
(Dahl 1931)
speciesimage
female Vulva
(Loksa 1969)
Further taxa

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1. Coelotes alpinus Polenec, 1972

speciesimage
male Pedipalp, retrolateral view
(Polenec 1972)
speciesimage
male Pedipalp, prolateral view
(Polenec 1972)
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male Pedipalp, ventral view
(Polenec 1972)
speciesimage
female Epigyne, ventral view
(Wang & Jäger 2010)
speciesimage
female Epigyne, ventral view
(Polenec 1972)
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female Vulva
(Polenec 1972)

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2. Coelotes arganoi Brignoli, 1978

speciesimage
female Epigyne, ventral view
(Brignoli 1978a)
speciesimage
female Epigyne, ventral view
(Wang & Zhu 2009)
speciesimage
female Vulva, dorsal view
(Brignoli 1978a)
speciesimage
female Vulva, dorsal view
(Wang & Zhu 2009)
speciesimage
female Eyes, dorsal view
(Wang & Zhu 2009)

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3. Coelotes coenobita Brignoli, 1978

speciesimage
female Epigyne, ventral view
(Brignoli 1978a)
speciesimage
female Epigyne, ventral view
(Wang & Zhu 2009)
speciesimage
female Vulva, dorsal view
(Brignoli 1978a)
speciesimage
female Vulva, dorsal view
(Wang & Zhu 2009)
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female Eyes, dorsal view
(Wang & Zhu 2009)

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4. Coelotes italicus Kritscher, 1956

speciesimage
male Pedipalp, retrolateral view
(De Blauwe 1973)
speciesimage
male Pedipalp, dorsal view
(De Blauwe 1973)
speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Tip of pedipalp, ventral view
(De Blauwe 1973)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Vulva
(De Blauwe 1973)

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5. Coelotes luculli Brignoli, 1978

speciesimage
female Epigyne, ventral view
(Brignoli 1978a)
speciesimage
female Vulva, dorsal view
(Brignoli 1978a)

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6. Coelotes osellai De Blauwe, 1973

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Vulva
(De Blauwe 1973)
speciesimage
female Vulva
(Maurer 1982a)

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7. Coelotes pabulator Simon, 1875

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(De Blauwe 1975)

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8. Coelotes phthisicus Brignoli, 1978

speciesimage
male Pedipalp, ventral view
(Danişman 2013b)
speciesimage
male Pedipalp, retrolateral view
(Danişman 2013b)
speciesimage
male Pedipalp, prolateral view
(Danişman 2013b)
speciesimage
male Habitus, dorsal view
(Danişman 2013b)
speciesimage
female Epigyne, ventral view
(Brignoli 1978a)
speciesimage
female Vulva, dorsal view
(Brignoli 1978a)
speciesimage
female Habitus, dorsal view
(Danişman 2013b)

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9. Coelotes pickardi O. P.-Cambridge, 1873

speciesimage
male Pedipalp, ventral view
(Isaia & Pantini 2009)
speciesimage
male Conductor
(Isaia & Pantini 2009)
speciesimage
female Epigyne
(De Blauwe 1975)
speciesimage
female Vulva
(De Blauwe 1975)

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10. Coelotes pickardi carpathensis Ovtchinnikov, 1999

speciesimage
male Pedipalp, ventral view
(Ovtchinnikov 1999)
speciesimage
male Pedipalp, retrolateral view
(Ovtchinnikov 1999)
speciesimage
female Epigyne, ventral view
(Ovtchinnikov 1999)
speciesimage
female Vulva, dorsal view
(Ovtchinnikov 1999)

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11. Coelotes pickardi pastor Simon, 1875

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Conductor
(Isaia & Pantini 2009)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
female Epigyne
(De Blauwe 1973)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(De Blauwe 1975)
speciesimage
female Vulva
(De Blauwe 1975)

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12. Coelotes pickardi tirolensis (Kulczyński, 1906)

speciesimage
male Conductor
(Isaia & Pantini 2009)
speciesimage
female Epigyne, ventral view
(Drensky 1942)

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13. Coelotes poleneci Wiehle, 1964

speciesimage
male Pedipalp, retrolateral view
(Wiehle 1964b)
speciesimage
male Pedipalp, prolateral view
(Wiehle 1964b)
speciesimage
male Bulbus, ventral view
(Wiehle 1964b)
speciesimage
female Epigyne, ventral view
(Wiehle 1964b)
speciesimage
female Vulva
(Wiehle 1964b)

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14. Coelotes poweri Simon, 1875

speciesimage
male Pedipalp, ventral view
(De Blauwe 1973)
speciesimage
male Pedipalp
(De Blauwe 1973)
speciesimage
male Patellar apophysis
(De Blauwe 1973)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)

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15. Coelotes rhododendri Brignoli, 1978

speciesimage
male Pedipalp, retrolateral view
(Brignoli 1978a)
speciesimage
male Pedipalp, prolateral view
(Brignoli 1978a)

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16. Coelotes rudolfi (Schenkel, 1925)

speciesimage
male Pedipalp, ventral view
(Maurer 1982a)
speciesimage
male Pedipalp
(Maurer 1982a)
speciesimage
male Pedipalp, dorsal view
(De Blauwe 1975)
speciesimage
male Pedipalp, retrolateral view
(De Blauwe 1975)
speciesimage
male Pedipalp, ventral view
(De Blauwe 1975)
speciesimage
male Conductor
(Isaia & Pantini 2009)
speciesimage
male Patella and tibia, dorsal view
(De Blauwe 1975)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Epigyne
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)
speciesimage
female Vulva
(Maurer 1982a)

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17. Coelotes sinopensis Danişman, Karanfil & Coşar, 2016

speciesimage
male Pedipalp, retrolateral view
(Danişman et al. 2016)
speciesimage
male Pedipalp, ventral view
(Danişman et al. 2016)
speciesimage
female Epigyne, ventral view
(Danişman et al. 2016)
speciesimage
female Epigyne, posterior view
(Danişman et al. 2016)
speciesimage
female Vulva, dorsal view
(Danişman et al. 2016)
speciesimage
female Habitus, dorsal view
(Danişman et al. 2016)

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18. Coelotes titaniacus Brignoli, 1977

speciesimage
female Epigyne, ventral view
(Brignoli 1977a)
speciesimage
female Vulva, dorsal view
(Brignoli 1977a)

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19. Coelotes turkestanicus Ovtchinnikov, 1999

speciesimage
male Pedipalp, retrolateral view
(Esyunin et al. 2007)
speciesimage
male Pedipalp, ventral view
(Esyunin et al. 2007)
speciesimage
female Epigyne, ventral view
(Esyunin et al. 2007)
speciesimage
female Vulva
(Esyunin et al. 2007)

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20. Coelotes vallei Brignoli, 1977

speciesimage
female Vulva, ventral view
(Brignoli 1977b)

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21. Coelotes vignai Brignoli, 1978

speciesimage
female Epigyne/vulva, ventral view
(Brignoli 1978a)
speciesimage
female Epigyne, ventral view
(Wang & Zhu 2009)
speciesimage
female Vulva, dorsal view
(Wang & Zhu 2009)
speciesimage
female Eyes, dorsal view
(Wang & Zhu 2009)