Gnaphosidae: Zelotes Gistel, 1848

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1
Males

  2  

- Females

  25  

2
(1)
Embolus long, whip-like, forming a loop

  3  

- Different

  5  

3
(2)
Body length < 2.5 mm

  4  

-

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Body length 4-7 mm; bulbus as fig.; opisthosoma with dark brown scuticula

Civizelotes caucasius (L. Koch, 1866)

female

4
(3)

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cp

Embolic base with 2 thorn-like apophyses; opisthosoma with scuticula

Civizelotes pygmaeus (Miller, 1943)

female

-

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cp

Different, as fig.; opisthosoma with weakly developed scuticula

Civizelotes gracilis (Canestrini, 1868)

female

5
(2)
Embolic division not or hardly segmented or structures covered, thus at most tip of embolus or a apophysis is visible

  6  

- Embolic division of bulbus more strongly segmented, more details visible

  11  

6
(5)

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Embolic division hardly segmented, more details visible

Zelotes exiguus (Müller & Schenkel, 1895)

female

- At least tip of embolus visible

  7  

7
(6)

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cp

Tibial apophysis smaller/equal tibia; opisthosoma with dark brown scuticula

Zelotes puritanus Chamberlin, 1922

female

- Tibial apophysis larger or equal tibia

  8  

8
(7)

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cp

Bulbus with wedge-like apophysis; body length 3-5 mm

Civizelotes civicus (Simon, 1878)

female

- Different

  9  

9
(8)

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cp

Embolus triangularly pointed; tibial apophysis short and triangular; prosoma anteriorly broader than in other Zelotes-species

Zelotes segrex (Simon, 1878)

female

- Embolus different; tibial apophysis long and slender

  10  

10
(9)

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cp

Embolus short and broad

Zelotes hermani (Chyzer, 1897)

female

-

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Embolus short and rod-like

Zelotes latreillei (Simon, 1878)

female

11
(5)
Embolic division with transverse keel

  12  

- Different

  22  

12
(11)

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cp

Prosoma yellow-brown to light orange; embolus short and stout; legs distally black-brown; keel weakly developed

Zelotes electus (C. L. Koch, 1839)

female

- Uniform colour

  13  

13
(12)
Embolus long, coiled or with lobe shaped apophysis

  14  

- Embolus different, mostly shorter

  16  

14
(13)

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cp

Embolus distally with lobe-shaped apophysis; tibial apophysis straight

Zelotes apricorum (L. Koch, 1876)

female

- Different

  15  

15
(14)

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cp

Embolus spirally coiled; keel with long, fine tip; tibial apophysis slightly S-shaped

Zelotes erebeus (Thorell, 1871)

female

-

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Embolus broad, bent; tibial apophysis slightly bent

Zelotes tenuis (L. Koch, 1866)

female

16
(13)

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cp

Tip of keel very long and thin, reaching margin of cymbium

Zelotes aeneus (Simon, 1878)

female

-

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cp

Further species

Zelotes zellensis Grimm, 1982

female

- Different

  17  

17
(16)

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cp

Bulbus anterior to keel with conspicuous apophysis

Zelotes atrocaeruleus (Simon, 1878)

female

- Bulbus without such apophysis

  18  

18
(17)

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cp

Embolus short and robust; keel partially covered by distal part of bulbus

Zelotes gallicus Simon, 1914

female

- Different

  19  

19
(18)

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cp

Embolus long and pointed; keel at inner side of bulbus more covered by distal part of bulbus

Zelotes clivicola (L. Koch, 1870)

female

-

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Similar to clivicola, tibial apophysis straight, longer, reaching to embolic base though

Zelotes devotus Grimm, 1982

female

- Different

  20  

20
(19)

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cp

Keel distally with stepped edge

Zelotes similis (Kulczyński, 1887)

female

- Different

  21  

21
(20)

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cp

Tip of keel bent; embolus short and stout

Zelotes aurantiacus Miller, 1967

female

-

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cp

Tip of keel straight; Embolus long and narrow

Zelotes subterraneus (C. L. Koch, 1833)

female

22
(11)

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cp

Tibial apophysis about twice as long as tibia

Zelotes longipes (L. Koch, 1866)

female

- Length of tibial apophysis ± length of tibia

  23  

23
(22)

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cp

Bulbus with several hook-shaped apophyses

Zelotes petrensis (C. L. Koch, 1839)

female

- Different

  24  

24
(23)

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cp

Bulbus distally with a fringy apophysis on the outer side; Embolus thin and straight

Zelotes oblongus (C. L. Koch, 1833)

female

-

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Fringy apophysis missing; embolus long and bent, with sickle-shaped apophysis

Zelotes talpinus (L. Koch, 1872)

female

25
(1)
Anterior epigynal margin with 2 ear-shaped protrusions, which are distinctly marked off from remaining epigyne

  26  

- Anterior epigynal margin without such distinctly marked off structures

  45  

26
(25)
Posterior epigynal margin with transverse cuticular fold or grooves

  27  

- Cuticular fold on posterior epigynal margin not interrupted

  31  

27
(26)

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cp

Several transverse, curved cuticular folds below the ear-shaped protrusion

Zelotes latreillei (Simon, 1878)

male

- Different

  28  

28
(27)

world

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cp

Anterior epigynal region with a transverse, bar-like protrusion

Civizelotes civicus (Simon, 1878)

male

- Different

  29  

29
(28)

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cp

Lateral cuticular folds parallel in the anterior epigynal region

Zelotes talpinus (L. Koch, 1872)

male

- Lateral cuticular folds distinctly converging anteriorly

  30  

30
(29)

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cp

Lateral cuticular folds bent or rounded

Zelotes longipes (L. Koch, 1866)

male

-

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cp

Lateral cuticular folds running straight, similar to longipes

Zelotes hermani (Chyzer, 1897)

male

31
(26)

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cp

Epigyne with 3 pairs of protrusions and S-shaped cuticular fold

Zelotes petrensis (C. L. Koch, 1839)

male

- Different

  32  

32
(31)

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cp

Epigyne hardly structured, only with semicircular groove

Zelotes segrex (Simon, 1878)

male

- Ear-shaped protrusions distinctly separated

  33  

33
(32)
2 separated cuticular ridges running from ear-shaped protrusions to posterior epigynal margin

  34  

- Different

  35  

34
(33)

world

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cp

Length of these cuticular ridges = distance of the ear-shaped structures to each other

Zelotes erebeus (Thorell, 1871)

male

-

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cp

Length of cuticular ridges much less than distance of the ear-shaped structures

Zelotes atrocaeruleus (Simon, 1878)

male

35
(33)

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cp

Epigyne apart of a semicircular cuticular fold without strongly sclerotised regions

Zelotes oblongus (C. L. Koch, 1833)

male

- Epigyne more strongly sclerotised; area between ear-shaped protrusions and posterior epigynal part less sclerotised

  36  

36
(35)
Epigyne with leaf-shaped protrusions of cuticular ridges

  37  

- Without such leaf-shaped protrusions

  38  

37
(36)

world

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cp

Cuticular fold pointed S-shaped, running back posteriorly

Zelotes aurantiacus Miller, 1967

male

-

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cp

Cuticular fold only slightly rounded

Zelotes clivicola (L. Koch, 1870)

male

38
(36)
Opening of the anterior cuticular fold narrow, about 1/3 - 1/5 of distance between ear-shaped protrusions

  39  

- Opening wide, greater or equal 1/2 of distance between ear-shaped protrusions

  40  

39
(38)

world

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cp

Anterior cuticular fold parallel to posterior cuticular fold

Zelotes apricorum (L. Koch, 1876)

male

-

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cp

Anterior cuticular fold transverse to posterior cuticular fold

Zelotes devotus Grimm, 1982

male

40
(38)

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cp

Lateral cuticular ridges approaching each other up to 1/5 of distance between ear-shaped protrusions

Zelotes zellensis Grimm, 1982

male

- Lateral cuticular ridges approaching about 1/2 - 3/4 of distance between ear-shaped protrusions

  41  

41
(40)

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cp

Lateral cuticular ridges ± straight-lined

Zelotes similis (Kulczyński, 1887)

male

- Lateral cuticular ridges distinctly curved

  42  

42
(41)
Lateral and median protrusions of posterior cuticular ridge forming an uniform, doubly S-shaped line

  43  

- Different

  44  

43
(42)

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cp

Epigyne and vulva as figs.

Zelotes subterraneus (C. L. Koch, 1833)

male

-

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cp

Epigyne distinctly less sclerotised, primarily in posterior region; anterior margin reduced to two semicircles

Zelotes aeneus (Simon, 1878)

male

44
(42)

world

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cp

Lateral protrusion of posterior cuticular ridge overlapping median protrusion posteriorly

Zelotes electus (C. L. Koch, 1839)

male

-

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Median protrusion of posterior cuticular ridge overlapping lateral protrusion posteriorly

Zelotes gallicus Simon, 1914

male

45
(25)
Epigyne rounded, ± uniformly sclerotised

  48  

- Different

  46  

46
(45)

world

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cp

Posterior epigynal margin with 3/4 circular, closed cuticular fold

Zelotes puritanus Chamberlin, 1922

male

- Different

  47  

47
(46)

world

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cp

Epigyne without longitudinal cuticular ridge, hardly sclerotised

Zelotes exiguus (Müller & Schenkel, 1895)

male

-

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cp

Epigyne only in anterior area sclerotised

Civizelotes caucasius (L. Koch, 1866)

male

-

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cp

Further species

Zelotes tenuis (L. Koch, 1866)

male

48
(45)

world

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cp

Cuticular ridge of epigyne roundish-rectangular

Civizelotes pygmaeus (Miller, 1943)

male

-

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cp

Cuticular ridge of epigyne straightly converging backwards

Civizelotes gracilis (Canestrini, 1868)

male

Further taxa

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1. Zelotes aerosus Charitonov, 1946

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2. Zelotes alpujarraensis Senglet, 2011

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3. Zelotes argoliensis (C. L. Koch, 1839)

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4. Zelotes azsheganovae Esyunin & Efimik, 1992

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5. Zelotes babunaensis (Drensky, 1929)

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6. Zelotes baeticus Senglet, 2011

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7. Zelotes balcanicus Deltshev, 2006

world

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8. Zelotes bimaculatus (C. L. Koch, 1837)

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9. Zelotes boluensis Wunderlich, 2011

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10. Zelotes calactinus Di Franco, 1989

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11. Zelotes callidus (Simon, 1878)

world

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12. Zelotes chaniaensis Senglet, 2011

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13. Zelotes cingarus (O. P.-Cambridge, 1874)

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14. Zelotes cordubensis Senglet, 2011

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15. Zelotes cornipalpus Melic, Silva & Barrientos, 2016

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16. Zelotes creticus (Kulczyński, 1903)

world

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17. Zelotes criniger Denis, 1937

world

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18. Zelotes cyanescens Simon, 1914

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19. Zelotes daidalus Chatzaki, 2003

world

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20. Zelotes denapes Platnick, 1993

world

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cp

21. Zelotes egregioides Senglet, 2011

world

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22. Zelotes egregius Simon, 1914

world

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23. Zelotes eugenei Kovblyuk, 2009

world

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cp

24. Zelotes fallax Tuneva & Esyunin, 2003

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25. Zelotes femellus (L. Koch, 1866)

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26. Zelotes flagellans (L. Koch, 1882)

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27. Zelotes flavimanus (C. L. Koch, 1839)

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28. Zelotes fulvaster (Simon, 1878)

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29. Zelotes fulvopilosus (Simon, 1878)

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30. Zelotes fuscorufus (Simon, 1878)

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31. Zelotes fuscus (Thorell, 1875)

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32. Zelotes fuzeta Wunderlich, 2011

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33. Zelotes harmeron Levy, 2009

world

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34. Zelotes helicoides Chatzaki, 2010

world

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35. Zelotes hispaliensis Senglet, 2011

world

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36. Zelotes khostensis Kovblyuk & Ponomarev, 2008

world

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cp

37. Zelotes laconicus Senglet, 2011

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cp

38. Zelotes laetus (O. P.-Cambridge, 1872)

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39. Zelotes lagrecai Di Franco, 1994

world

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cp

40. Zelotes longestylus Simon, 1914

world

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cp

41. Zelotes maccaricus Di Franco, 1998

world

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cp

42. Zelotes manius (Simon, 1878)

world

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cp

43. Zelotes messinai Di Franco, 1995

world

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cp

44. Zelotes metellus (Roewer, 1928)

world

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cp

45. Zelotes minous Chatzaki, 2003

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cp

46. Zelotes mundus (Kulczyński, 1897)

world

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cp

47. Zelotes murcidus Simon, 1914

world

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cp

48. Zelotes occidentalis Melic, 2014

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cp

49. Zelotes occultus Tuneva & Esyunin, 2003

world

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cp

50. Zelotes olympi (Kulczyński, 1903)

world

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cp

51. Zelotes orenburgensis Tuneva & Esyunin, 2003

world

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cp

52. Zelotes paroculus Simon, 1914

world

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cp

53. Zelotes pediculatoides Senglet, 2011

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cp

54. Zelotes pediculatus Marinaro, 1967

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cp

55. Zelotes plumiger (L. Koch, 1882)

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cp

56. Zelotes pluridentatus Marinaro, 1967

world

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cp

57. Zelotes potanini Schenkel, 1963

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cp

58. Zelotes prishutovae Ponomarev & Tsvetkov, 2006

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cp

59. Zelotes pseudogallicus Ponomarev, 2007

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cp

60. Zelotes pyrenaeus Di Franco & Blick, 2003

world

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cp

61. Zelotes reimoseri Roewer, 1951

world

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cp

62. Zelotes rinske van Helsdingen, 2012

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cp

63. Zelotes rufi Esyunin & Efimik, 1997

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64. Zelotes sardus (Canestrini, 1873)

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65. Zelotes scrutatus (O. P.-Cambridge, 1872)

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66. Zelotes semibadius (L. Koch, 1878)

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67. Zelotes serratus Wunderlich, 2011

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68. Zelotes spadix (L. Koch, 1866)

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69. Zelotes strandi (Nosek, 1905)

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70. Zelotes thorelli Simon, 1914

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71. Zelotes turcicus Wunderlich, 2011

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72. Zelotes uronesae Melic, 2014

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cp

73. Zelotes vespertinus (Thorell, 1875)

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74. Zelotes wallacei Melic, Silva & Barrientos, 2016

world75. Zelotes caprearum (Pavesi, 1875)
world76. Zelotes distinctissimus Caporiacco, 1929
world77. Zelotes graecus (L. Koch, 1867)
world78. Zelotes hirtus (Thorell, 1875)
world79. Zelotes insulanus (L. Koch, 1867)
world80. Zelotes insulanus (2) Dalmas, 1922
world81. Zelotes siculus (Simon, 1878)
world82. Zelotes tristis (Thorell, 1871)